Thalassemydidae
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''Thalassemys'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
thalassochelydia Thalassochelydia is a clade of extinct marine turtles from the Late Jurassic and earliest Cretaceous of Europe and South America. The group is defined as including ''Eurysternum'', ''Plesiochelys'' and ''Thalassemys'' to the exclusion of ''Pelome ...
n
turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked tu ...
from the
Late Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 163.5 ± 1.0 to 145.0 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
of western and central
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
. While the genus was originally named by Rütimeyer in 1859 for a large
carapace A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tor ...
and other associated fragments from the late
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 157.3 ± 1.0 Ma and 152.1 ± 0.9 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...
of the
Reuchenette Formation The Reuchenette Formation is a Jurassic geologic formation in Switzerland. It is Kimmeridgian in age and predominantly consists of well stratified limestone, with lithology variable both laterally and stratigraphically including wackestones, pack ...
of
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
, although the taxon was not validly named until 1873 when Rütimeyer designated the type species ''T. hugii''. Rütimeyer also named ''T. gresslyi'' from the Reunchenette Formation in the same paper as ''T. hugii'', but it cannot be differentiated from the type material of ''T. hugii'' and is therefore a junior synonym. A large assemblage of shell and postcranial material from the Reunchenette was named as a species of ''
Eurysternum ''Eurysternum'' is an extinct genus of thalassochelydian turtle. Its type species is ''Eurysternum wagleri'', the holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species ...
'', ''E. ignoratum'', by Bräm in
1965 Events January–February * January 14 – The Prime Minister of Northern Ireland and the Taoiseach of the Republic of Ireland meet for the first time in 43 years. * January 20 ** Lyndon B. Johnson is Second inauguration of Lyndo ...
. While originally distinguished based on the presence of fontanelles on the plastron, the feature was later identified on ''T. hugii'' and ''E. ignoratum'' was designated a junior synonym. Additional material from the
Kimmeridge Clay The Kimmeridge Clay is a sedimentary deposit of fossiliferous marine clay which is of Late Jurassic to lowermost Cretaceous age and occurs in southern and eastern England and in the North Sea. This rock formation is the major source rock for North ...
of the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...
has also been referred to ''T. hugii''. The second valid species of ''Thalassemys'' is ''T. marina'', named in 1903 by Fraas for a partial carapace and half a plastron from the late Kimmeridgian deposits of Schnaitheim,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
. It was suggested to be a species of the related taxon ''
Eurysternum ''Eurysternum'' is an extinct genus of thalassochelydian turtle. Its type species is ''Eurysternum wagleri'', the holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species ...
'' by Maisch in 2001, who referred additional material from Schnaitheim to the species. In 2003, it was suggested that ''T. marina'' was a synonym of the related turtle ''
Palaeomedusa ''Palaeomedusa testa'' is an extinct species of thalassochelydian turtle from the Tithonian of the Late Jurassic (145.5 to 150.8 million years ago). It was first described by the German palaeontologist Christian Erich Hermann von Meyer in 1860.F ...
'', which was based on an unknown but restored region of the original carapace. ''Thalassemys heusseri'' was validly named in 1924 by Oertel for a partial carapace from the Kimmeridgian or
Tithonian In the geological timescale, the Tithonian is the latest age of the Late Jurassic Epoch and the uppermost stage of the Upper Jurassic Series. It spans the time between 152.1 ± 4 Ma and 145.0 ± 4 Ma (million years ago). It is preceded by the K ...
of
Holzen Holzen is a municipality in the district of Holzminden, in Lower Saxony, Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, a ...
, Germany, a different spelling from the invalid name "T. heuseri" he used previously in
1815 Events January * January 2 – Lord Byron marries Anna Isabella Milbanke in Seaham, county of Durham, England. * January 3 – Austria, Britain, and Bourbon-restored France form a secret defensive alliance treaty against Prussi ...
. As the type specimen was lost, the species cannot be distinguished from others of the genus, and is therefore a '' nomen dubium''. In addition to ''E. ignoratum'', Bräm also named the species ''Thalassemys moseri'' in 1965 for a partial carapace from the Reunchenette Formation. An additional shell and also a skull were referred to the taxon in 1980 by Rieppel, who suggested that the species should be included in ''
Plesiochelys ''Plesiochelys'' is a genus of late Jurassic European and Asian turtle. The type species is ''Plesiochelys etalloni''. Fossil records have discovered ''Plesiochelys bigleri'' and ''Plesiochelys etalloni'' from the Kimmeridge Clay of England and ...
'' rather than ''Thalassemys''. This skull was given the name '' Jurassichelon'' in 2015, leaving the species ''T. moseri'', based on other material, in an uncertain position. Further assessment of the material by Anquetin and colleagues in 2017 concluded that ''T. moseri'' was distinct from the species ''J. oleronensis'' named for the skull, but was representative of the same genus. ''Thalassemys bruntrutana'' was described in 2015 by Püntener and colleagues for an almost complete carapace and plastron from a lower member of the Reunchenette Formation than other species, with referred material also from the British
Kimmeridge Clay The Kimmeridge Clay is a sedimentary deposit of fossiliferous marine clay which is of Late Jurassic to lowermost Cretaceous age and occurs in southern and eastern England and in the North Sea. This rock formation is the major source rock for North ...
of the
Isle of Purbeck The Isle of Purbeck is a peninsula in Dorset, England. It is bordered by water on three sides: the English Channel to the south and east, where steep cliffs fall to the sea; and by the marshy lands of the River Frome and Poole Harbour to the no ...
. While the genus ''Thalassemys'' was placed in the family Thalassemydidae, it represents the only definitive member of the family due to the uncertain relationships of other members of
Thalassochelydia Thalassochelydia is a clade of extinct marine turtles from the Late Jurassic and earliest Cretaceous of Europe and South America. The group is defined as including ''Eurysternum'', ''Plesiochelys'' and ''Thalassemys'' to the exclusion of ''Pelome ...
, primarily due to the lack of cranial material in ''Thalassemys''.


References

* {{Taxonbar, from=Q7709531 Thalassochelydia Prehistoric turtle genera Kimmeridgian genera Late Jurassic turtles Late Jurassic reptiles of Europe Jurassic England Fossils of England Jurassic Germany Fossils of Germany Jurassic Switzerland Fossils of Switzerland Fossil taxa described in 1873