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''Podocarpus totara'' (; from the Maori-language ; the spelling "totara" is also common in English) is a species of
podocarp Podocarpaceae is a large family of mainly Southern Hemisphere conifers, known in English as podocarps, comprising about 156 species of evergreen trees and shrubs.James E. Eckenwalder. 2009. ''Conifers of the World''. Portland, Oregon: Timber Pre ...
tree
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
. It grows throughout the
North Island The North Island, also officially named Te Ika-a-Māui, is one of the two main islands of New Zealand, separated from the larger but much less populous South Island by the Cook Strait. The island's area is , making it the world's 14th-largest ...
and northeastern
South Island The South Island, also officially named , is the larger of the two major islands of New Zealand in surface area, the other being the smaller but more populous North Island. It is bordered to the north by Cook Strait, to the west by the Tasman ...
in lowland, montane and lower subalpine forest at elevations of up to 600 m. Tōtara is commonly found in lowland areas where the soil is fertile and well drained.


Description

The tōtara is a medium to large
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
, which grows slowly to around 20 to 25 m, exceptionally to 35 m; it is noted for its longevity and the great girth of its trunk. The bark peels off in papery flakes, with a purplish to golden brown hue. The sharp, dull-green, needle-like
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are stiff and leathery, 2 cm long. This plant produces highly modified
cones A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex. A cone is formed by a set of line segments, half-lines, or lines conn ...
with two to four fused, fleshy, berry-like, juicy scales, bright red when mature. The cone contains one or two rounded
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
s at the apex of the scales. The largest known living tōtara, the Pouakani Tree, near Pureora in the central North Island, is over 35 m tall and nearly 4 m in trunk diameter at breast height. Bushmen discovered it in 1950. Other large trees are known in this area, while Whirinaki Forest, to the east, but also on deep recent volcanic soils, has groves of very tall tōtara (over 40 m in height). Tōtara is often found regenerating on farmland, as it is not eaten by livestock.


Varieties

The two varieties of tōtara are: * ''P. t.'' var. ''totara'' * ''P. t.'' var. ''waihoensis''


Related trees

In a classic example of
Antarctic flora Antarctic flora are a distinct community of vascular plants which evolved millions of years ago on the supercontinent of Gondwana. Presently, species of Antarctica flora reside on several now separated areas of the Southern Hemisphere, includin ...
species-pair the tōtara is very closely related to ''
Podocarpus nubigenus ''Podocarpus nubigenus'' (also known as ''P. nubigena'') is a species of Podocarpus, podocarp, endemic (ecology), endemic to the Valdivian temperate rain forests of southern Chile and adjacent southwestern Argentina. It is the southernmost podoca ...
'' from
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, to the extent that if planted together, they are very difficult to distinguish. The best distinction is the grey-green tone of the leaves, compared to the slightly brighter green of ''P. nubigenus''.


Cultivation

Tōtara grows easily from fresh seed and cuttings. It has been planted in the United Kingdom as far north as Inverewe,
Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the ...
. Several cultivars for garden use have been introduced. These include 'Albany Gold' and 'Aurea', both have yellow 'gold' foliage that darkens in winter; 'Pendula', which has a weeping growth habit that is especially pronounced in young plants; 'Silver Falls', also pendulous but with cream-edged foliage; and 'Matapouri Blue', which has a conical form and glaucous foliage.


Human use

The
wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin th ...
is hard, straight-grained, and very resistant to rot, especially its
heartwood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin tha ...
. Due to its durability, tōtara wood was often used for fence posts, floor pilings, and railway sleepers. It is also prized for its carving properties, and was the primary wood used in
Māori Māori or Maori can refer to: Relating to the Māori people * Māori people of New Zealand, or members of that group * Māori language, the language of the Māori people of New Zealand * Māori culture * Cook Islanders, the Māori people of the C ...
carving. It was the primary wood used to make ''waka'' in traditional Maori boat building due to its relatively light weight (about 25% lighter than
kauri ''Agathis'', commonly known as kauri or dammara, is a genus of 22 species of evergreen tree. The genus is part of the ancient conifer family Araucariaceae, a group once widespread during the Jurassic and Cretaceous periods, but now largely res ...
), long, straight lengths, and natural oils in the wood that help prevent rotting. Tōtara could be drilled with chert points to make holes near the edges of the timber without splitting. In larger tōtara ''waka'', three or more sections were laced together with flax rope. A tōtara ''waka'' took at least a year to make using stone adzes. Bark from tōtara is used to cover and protect traditional
pōhā Pōhā are traditional Māori bags made from southern bull kelp, which are used to carry and store food and fresh water, to propagate live shellfish, and to make clothing and equipment for sports. Pōhā are especially associated with Ngāi Tah ...
bags.


Gallery

image:PodocarpusTotara.jpg, Receptacle and seed of tōtara image:PodocarpusTotaraBark.jpg, The reddish-grey bark of the tōtara is thick, corky, furrowed and stringy


References


External links

* . * New Zealand Plant Conservation Network, UR
''Podocarpus totara'' var. ''totara''
Accessed 2010-10-03. * New Zealand Plant Conservation Network, UR
''Podocarpus totara'' var. ''waihoensis''
Accessed 2010-10-03. {{Taxonbar, from=Q164276
totara ''Podocarpus totara'' (; from the Maori-language ; the spelling "totara" is also common in English) is a species of podocarp tree endemic to New Zealand. It grows throughout the North Island and northeastern South Island in lowland, montane and ...
Trees of New Zealand Endemic flora of New Zealand