''Typhlatya'' is a genus of

shrimp Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are refer ...

in the family Atyidae. These are small,

stygobitic Stygofauna are any fauna that live in groundwater systems or aquifers, such as caves, fissures and vugs. Stygofauna and troglofauna are the two types of subterranean fauna (based on life-history). Both are associated with subterranean environments ...

shrimp found in the West

Mediterranean The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the e ...

region (France and Spain), Caribbean region (the

Antilles The Antilles (; gcf, label=Antillean Creole, Antiy; es, Antillas; french: Antilles; nl, Antillen; ht, Antiy; pap, Antias; Jamaican Patois: ''Antiliiz'') is an archipelago bordered by the Caribbean Sea to the south and west, the Gulf of Mex ...

Bahamas The Bahamas (), officially the Commonwealth of The Bahamas, is an island country within the Lucayan Archipelago of the West Indies in the Atlantic Ocean, North Atlantic. It takes up 97% of the Lucayan Archipelago's land area and is home to ...

and Yucatán),

Ascension Island Ascension Island is an isolated volcanic island, 7°56′ south of the Equator in the South Atlantic Ocean. It is about from the coast of Africa and from the coast of South America. It is governed as part of the British Overseas Territory o ...

and the Galápagos, although the individual species often have very small ranges. Species in this genus are found in salt,

brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...

and fresh waters, mostly in

anchialine An anchialine system (, from Greek ''ankhialos'', "near the sea") is a landlocked body of water with a subterranean connection to the ocean. Depending on its formation, these systems can exist in one of two primary forms: pools or caves. The primary ...

habitats and none in the open sea.

Taxonomy, species and conservation

There are currently 17 recognized species, of which one is listed by the

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

as critically endangered (marked "CR" below), and six are considered vulnerable (marked "VU").

Phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...

studies indicate that the genus as presently defined is

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

and needs to be redefined, as ''T. galapagensis'' clusters with the widespread '' Antecaridina'' and ''T. monae'' likely with the Australian ''

Stygiocaris ''Stygiocaris'' is a genus of shrimp from caves in the North West Cape and Barrow Island (Western Australia), Barrow Island, Western Australia. ''Stygiocaris'' is highly troglobite, troglobitic, being transparency and translucency, transparent, ...

''. On a higher level, these three genera, together with '' Halocaridina'' and '' Halocaridinides'', form a group. In addition to the species below, ''"Typhlatya" jusbaschjani'' (Georgia) and ''"Typhlatya" pretneri'' (Hercegovina) have been listed in this genus, but the former is a ''

species inquirenda In biological classification, a ''species inquirenda'' is a species of doubtful identity requiring further investigation. The use of the term in English-language biological literature dates back to at least the early nineteenth century. The term t ...

'' and recent authorities generally include the latter in ''

Troglocaris ''Troglocaris'' is a genus of freshwater shrimp in the family Atyidae. These stygobitic, whitish and eyeless shrimp are found in Southern Europe (Dinaric Alps and West Caucasus). Although locally very common, the small ranges of the individual s ...

''. *''

Typhlatya arfeae ''Typhlatya'' is a genus of shrimp in the family Atyidae. These are small, stygobitic shrimp found in the West Mediterranean region (France and Spain), Caribbean region (the Antilles, Bahamas and Yucatán), Ascension Island and the Galápagos ...

'' Jaume & Bréhier, 2005 – France *'' Typhlatya campecheae'' H. H. Hobbs III & H. H. Hobbs Jr., 1976 – Yucatán Peninsula *'' Typhlatya consobrina'' Botoşăneanu & Holthuis, 1970 – Cuba Status iucn VU icon

*'' Typhlatya dzilamensis'' Alvarez, Iliffe & Villalobos, 2005 – Yucatán Peninsula *'' Typhlatya elenae'' Juarrero, 1994 – Cuba Status iucn VU icon

*'' Typhlatya galapagensis'' Monod & Cals, 1970 – Galápagos Islands *'' Typhlatya garciadebrasi'' Juarrero de Varona & Ortiz, 2000 – Cuba Status iucn VU icon

*'' Typhlatya garciai'' Chace, 1942 – Cuba and possibly Caicos Island Status iucn VU icon

*'' Typhlatya iliffei'' C. W. J. Hart & Manning, 1981 – Bermuda Status iucn CR icon

*'' Typhlatya kakuki'' Alvarez, Iliffe & Villalobos, 2005 – Bahamas *'' Typhlatya miravetensis'' Sanz & Platvoet, 1995 – Spain Status iucn VU icon

*'' Typhlatya mitchelli'' H. H. Hobbs III & H. H. Hobbs Jr., 1976 – Yucatán Peninsula *''

Typhlatya monae ''Typhlatya monae'' is a species of basket shrimp in the family Atyidae. It is found in the Caribbean and Africa. The IUCN conservation status of ''Typhlatya monae'' is "LC", least concern, with no immediate threat to the species' survival. The ...

'' Chace, 1954 – widely across Caribbean islands *'' Typhlatya pearsei'' Creaser, 1936 – Yucatán Peninsula *''

Typhlatya rogersi ''Typhlatya rogersi'' is a species of basket shrimp in the family Atyidae. It is found in South America and Africa. References Further reading * Atyidae Articles created by Qbugbot Crustaceans described in 1972 {{Caridea-stub ...

'' Chace & Manning, 1972 – Ascension Island *'' Typhlatya taina'' Estrada & Gómez, 1987 – Cuba Status iucn VU icon

*'' Typhlatya utilaensis'' Alvarez, Iliffe & Villalobos, 2005 – Utila Island