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''Typha'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of about 30
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
monocotyledon Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one embryonic leaf, or cotyledon. They constitute one of ...
ous
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s in the family
Typhaceae The Typhaceae () are a family of flowering plants, sometimes called the cattail family. The botanical name for the family has been recognized by most taxonomists. Description Members can be recognized as large marsh herbs with alternate two-rank ...
. These plants have a variety of common names, in
British English British English (BrE, en-GB, or BE) is, according to Lexico, Oxford Dictionaries, "English language, English as used in Great Britain, as distinct from that used elsewhere". More narrowly, it can refer specifically to the English language in ...
as bulrush or reedmace, in
American English American English, sometimes called United States English or U.S. English, is the set of variety (linguistics), varieties of the English language native to the United States. English is the Languages of the United States, most widely spoken lan ...
as reed, cattail, or punks, in
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
as cumbungi or bulrush, in
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
as bulrush or cattail, and in
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
as raupo. Other taxa of plants may be known as
bulrush Bulrush is a vernacular name for several large wetland grass-like plants *Sedge family (Cyperaceae): **''Cyperus'' **'' Scirpus'' **''Blysmus'' **''Bolboschoenus'' **'' Scirpoides'' **'' Isolepis'' **'' Schoenoplectus'' **'' Trichophorum'' * T ...
, including some
sedges The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus ''Carex'' wit ...
in ''
Scirpus ''Scirpus'' is a genus of grass-like species in the sedge family Cyperaceae many with the common names club-rush, wood club-rush or bulrush (see also bulrush for other plant genera so-named). They mostly inhabit wetlands and damp locations. Taxo ...
'' and related genera. The genus is largely distributed in the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
, where it is found in a variety of
wetland A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The ...
habitats. The rhizomes are edible. Evidence of preserved starch grains on
grinding stone Millstones or mill stones are stones used in gristmills, for grinding wheat or other grains. They are sometimes referred to as grindstones or grinding stones. Millstones come in pairs: a convex stationary base known as the ''bedstone'' and ...
s suggests they were already eaten in Europe 30,000 years ago.


Description

''Typha'' are aquatic or semi-aquatic, rhizomatous, herbaceous
perennial plant A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
s. The leaves are
glabrous Glabrousness (from the Latin '' glaber'' meaning "bald", "hairless", "shaved", "smooth") is the technical term for a lack of hair, down, setae, trichomes or other such covering. A glabrous surface may be a natural characteristic of all or part of ...
(hairless), linear, alternate and mostly basal on a simple, jointless stem that bears the flowering spikes. The plants are monoecious, with
unisexual Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproducti ...
flowers that develop in dense racemes. The numerous male flowers form a narrow spike at the top of the vertical stem. Each male (staminate) flower is reduced to a pair of
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s and hairs, and withers once the
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
is shed. Large numbers of tiny female flowers form a dense,
sausage A sausage is a type of meat product usually made from ground meat—often pork, beef, or poultry—along with salt, spices and other flavourings. Other ingredients, such as grains or breadcrumbs may be included as fillers or extenders. ...
-shaped spike on the stem below the male spike. In larger species this can be up to long and thick. The seeds are minute, long, and attached to fine hairs. When ripe, the heads disintegrate into a cottony fluff from which the seeds disperse by wind.


General ecology

''Typha'' are often among the first wetland plants to colonize areas of newly exposed wet mud, with their abundant wind-dispersed seeds. Buried seeds can survive in the soil for long periods of time. They
germinate Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, fer ...
best with sunlight and fluctuating temperatures, which is typical of many wetland plants that regenerate on mud flats. The plants also spread by rhizomes, forming large, interconnected stands. ''Typha'' are considered to be dominant competitors in wetlands in many areas, and they often exclude other plants with their dense canopy. In the bays of the
Great Lakes The Great Lakes, also called the Great Lakes of North America, are a series of large interconnected freshwater lakes in the mid-east region of North America that connect to the Atlantic Ocean via the Saint Lawrence River. There are five lak ...
, for example, they are among the most abundant wetland plants. Different species of cattails are adapted to different water depths. Well-developed
aerenchyma Aerenchyma or aeriferous parenchyma or lacunae, is a modification of the parenchyma to form a spongy tissue that creates spaces or air channels in the leaves, stems and roots of some plants, which allows exchange of gases between the shoot and th ...
make the plants tolerant of submersion. Even the dead stalks are capable of transmitting oxygen to the rooting zone. Although ''Typha'' are native wetland plants, they can be aggressive in their competition with other native species. They have been problematic in many regions in North America, from the Great Lakes to the
Everglades The Everglades is a natural region of tropical climate, tropical wetlands in the southern portion of the U.S. state of Florida, comprising the southern half of a large drainage basin within the Neotropical realm. The system begins near Orland ...
. Native sedges are displaced and wet meadows shrink, likely as a response to altered hydrology of the wetlands and increased nutrient levels. An introduced or hybrid species may be contributing to the problem. Control is difficult. The most successful strategy appears to be mowing or burning to remove the aerenchymous stalks, followed by prolonged flooding. It may be more important to prevent invasion by preserving water level fluctuations, including periods of drought, and to maintain infertile conditions. ''Typha'' are frequently eaten by wetland mammals such as muskrats, which also use them to construct feeding platforms and dens, thereby also providing nesting and resting places for waterfowl.


Accepted species and natural hybrids

The following names are currently accepted: The most widespread species is ''Typha latifolia'', which is distributed across the entire temperate northern hemisphere. It has also been introduced to Australia. ''T. angustifolia'' is nearly as widespread, but does not extend as far north; it may be introduced and invasive in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
. ''T. domingensis'' has a more southern
American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...
distribution, and it occurs in Australia. ''T. orientalis'' is widespread in Asia, Australia, and New Zealand. ''T. laxmannii'', ''T. minima'', and ''T. shuttleworthii'' are largely restricted to
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
and southern
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
.


Uses


Culinary

Many parts of the ''Typha'' plant are edible to humans. Before the plant flowers, the tender inside of the
shoot In botany, a plant shoot consists of any plant stem together with its appendages, leaves and lateral buds, flowering stems, and flower buds. The new growth from seed germination that grows upward is a shoot where leaves will develop. In the sp ...
s can be squeezed out and eaten raw or cooked. The starchy rhizomes are nutritious with a protein content comparable to that of maize or rice. They can be processed into a
flour Flour is a powder made by grinding raw grains, roots, beans, nuts, or seeds. Flours are used to make many different foods. Cereal flour, particularly wheat flour, is the main ingredient of bread, which is a staple food for many culture ...
with 266
kcal The calorie is a unit of energy. For historical reasons, two main definitions of "calorie" are in wide use. The large calorie, food calorie, or kilogram calorie was originally defined as the amount of heat needed to raise the temperature of o ...
per 100 grams, and are most often harvested from late autumn to early spring. They are fibrous, and the starch must be scraped or sucked from the tough fibers. Baby shoots emerging from the rhizomes, which are sometimes subterranean, can be picked and eaten raw. Also underground is a
carbohydrate In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may or ma ...
lump which can be peeled and eaten raw or cooked like a
potato The potato is a starchy food, a tuber of the plant ''Solanum tuberosum'' and is a root vegetable native to the Americas. The plant is a perennial in the nightshade family Solanaceae. Wild potato species can be found from the southern Unit ...
. The plant is one championed by survival experts because various parts can be eaten throughout the year. Plants growing in polluted water can accumulate
lead Lead is a chemical element with the symbol Pb (from the Latin ) and atomic number 82. It is a heavy metal that is denser than most common materials. Lead is soft and malleable, and also has a relatively low melting point. When freshly cu ...
and
pesticide Pesticides are substances that are meant to control pests. This includes herbicide, insecticide, nematicide, molluscicide, piscicide, avicide, rodenticide, bactericide, insect repellent, animal repellent, microbicide, fungicide, and lampri ...
residues in their rhizomes, and these should not be eaten. The rind of young stems can be peeled off, and the tender white heart inside can be eaten raw or boiled and eaten like
asparagus Asparagus, or garden asparagus, folk name sparrow grass, scientific name ''Asparagus officinalis'', is a perennial flowering plant species in the genus '' Asparagus''. Its young shoots are used as a spring vegetable. It was once classified in ...
. This food has been popular among the
Cossacks The Cossacks , es, cosaco , et, Kasakad, cazacii , fi, Kasakat, cazacii , french: cosaques , hu, kozákok, cazacii , it, cosacchi , orv, коза́ки, pl, Kozacy , pt, cossacos , ro, cazaci , russian: казаки́ or ...
in Russia, and has been called "Cossack asparagus". The leaf bases can be eaten raw or cooked, especially in late spring when they are young and tender. In early summer the sheath can be removed from the developing green flower spike, which can then be boiled and eaten like corn on the cob. In mid-summer when the male flowers are mature, the
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
can be collected and used as a flour supplement or thickener.


Agriculture

The seeds have a high
linoleic acid Linoleic acid (LA) is an organic compound with the formula COOH(CH2)7CH=CHCH2CH=CH(CH2)4CH3. Both alkene groups are cis-trans isomerism, ''cis''. It is a fatty acid sometimes denoted 18:2 (n-6) or 18:2 ''cis''-9,12. A linoleate is a salt (chem ...
content and can be used to feed cattle and chickens. They can also be found in African countries like Ghana. Harvesting cattail removes nutrients from the wetland that would otherwise return via the decomposition of decaying plant matter. Floating mats of cattails remove nutrients from eutrophied bodies of freshwater.


Building material

For local native tribes around
Lake Titicaca Lake Titicaca (; es, Lago Titicaca ; qu, Titiqaqa Qucha) is a large freshwater lake in the Andes mountains on the border of Bolivia and Peru. It is often called the highest navigable lake in the world. By volume of water and by surface area, ...
in
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
and
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
, ''Typha'' were among the most important plants and every part of the plant had multiple uses. For example, they were used to construct
raft A raft is any flat structure for support or transportation over water. It is usually of basic design, characterized by the absence of a hull. Rafts are usually kept afloat by using any combination of buoyant materials such as wood, sealed barrels ...
s and other boats. During World War II, the United States Navy used the down of ''Typha'' as a substitute for kapok in
life vests A personal flotation device (PFD; also referred to as a life jacket, life preserver, life belt, Mae West, life vest, life saver, cork jacket, buoyancy aid or flotation suit) is a flotation device in the form of a vest or suite that is worn by a ...
and aviation jackets. Tests showed that even after 100 hours of submersion, the buoyancy was still effective. ''Typha'' are used as
thermal insulation Thermal insulation is the reduction of heat transfer (i.e., the transfer of thermal energy between objects of differing temperature) between objects in thermal contact or in range of radiative influence. Thermal insulation can be achieved with ...
in buildings as an organic alternative to conventional insulating materials such as glass wool or stone wool.


Paper

''Typha'' stems and leaves can be used to make
paper Paper is a thin sheet material produced by mechanically or chemically processing cellulose fibres derived from wood, rags, grasses or other vegetable sources in water, draining the water through fine mesh leaving the fibre evenly distributed ...
. It is strong with a heavy texture and it is hard to bleach, so it is not suitable for industrial production of graphical paper. In 1853, considerable amounts of cattail paper were produced in New York, due to a shortage of raw materials. In 1948, French scientists tested methods for annual harvesting of the leaves. Because of the high cost, these methods were abandoned and no further research was done. Today ''Typha'' is used to make decorative paper.


Fiber

Fibers up to 4 meters long can be obtained from the stems when they are treated mechanically or chemically with sodium hydroxide. The stem fibers resemble
jute Jute is a long, soft, shiny bast fiber that can be spun into coarse, strong threads. It is produced from flowering plants in the genus ''Corchorus'', which is in the mallow family Malvaceae. The primary source of the fiber is ''Corchorus olit ...
and can be used to produce raw textiles. The leaf fibers can be used as an alternative to
cotton Cotton is a soft, fluffy staple fiber that grows in a boll, or protective case, around the seeds of the cotton plants of the genus ''Gossypium'' in the mallow family Malvaceae. The fiber is almost pure cellulose, and can contain minor perce ...
and linen in clothing. The yield of leaf fiber is 30 to 40 percent and ''Typha glauca'' can produce 7 to 10 tons per hectare annually.


Biofuel

''Typha'' can be used as a source of starch to produce
ethanol Ethanol (abbr. EtOH; also called ethyl alcohol, grain alcohol, drinking alcohol, or simply alcohol) is an organic compound. It is an Alcohol (chemistry), alcohol with the chemical formula . Its formula can be also written as or (an ethyl ...
. Because of their high productivity in northern latitudes, ''Typha'' are considered to be a bioenergy crop.


Other

The seed hairs were used by some
indigenous peoples of the Americas The Indigenous peoples of the Americas are the inhabitants of the Americas before the arrival of the European settlers in the 15th century, and the ethnic groups who now identify themselves with those peoples. Many Indigenous peoples of the A ...
as
tinder Tinder is easily combustible material used to start a fire. Tinder is a finely divided, open material which will begin to glow under a shower of sparks. Air is gently wafted over the glowing tinder until it bursts into flame. The flaming tinder i ...
for starting fires. Some tribes also used ''Typha'' down to line moccasins, and for bedding, diapers, baby powder, and
cradleboard Cradleboards (, se, gietkka, sms, ǩiõtkâm, smn, kietkâm, sje, gietkam) are traditional protective baby-carriers used by many indigenous cultures in North America and throughout northern Scandinavia amongst the Sámi. There are a variety ...
s. One Native American word for ''Typha'' meant "fruit for papoose's bed". ''Typha'' down is still used in some areas to stuff clothing items and pillows. ''Typha'' can be dipped in wax or fat and then lit as a candle, the stem serving as a wick. Without the use of wax or fat it will smolder slowly, somewhat like
incense Incense is aromatic biotic material that releases fragrant smoke when burnt. The term is used for either the material or the aroma. Incense is used for aesthetic reasons, religious worship, aromatherapy, meditation, and ceremony. It may also b ...
, and may repel insects. The flower stalks can be made into
chopsticks Chopsticks ( or ; Pinyin: ''kuaizi'' or ''zhu'') are shaped pairs of equal-length sticks of Chinese origin that have been used as kitchen and eating utensils in most of East and Southeast Asia for over three millennia. They are held in the do ...
. The leaves can be treated to weave into baskets, mats, or sandals. The rushes are harvested and the leaves often dried for later use in chair seats. Re-wetted, the leaves are twisted and wrapped around the chair rungs to form a densely woven seat that is then stuffed (usually with the left over rush). Small-scale experiments have indicated that ''Typha'' are able to remove
arsenic Arsenic is a chemical element with the symbol As and atomic number 33. Arsenic occurs in many minerals, usually in combination with sulfur and metals, but also as a pure elemental crystal. Arsenic is a metalloid. It has various allotropes, but ...
from drinking water. The boiled rootstocks have been used as a
diuretic A diuretic () is any substance that promotes diuresis, the increased production of urine. This includes forced diuresis. A diuretic tablet is sometimes colloquially called a water tablet. There are several categories of diuretics. All diuretics in ...
for increasing urination, or mashed to make a jelly-like paste for sores, boils, wounds, burns, scabs, and smallpox pustules. Cattail pollen is used as a banker source of food for predatory insects and mites (such as '' Amblyseius swirskii'') in greenhouses. The cattail, or, as it is commonly referred to in the American Midwest, the sausage tail, has been the subject of multiple artist renditions, gaining popularity in the mid-twentieth century. The term, sausage tail, derives from the similarity that cattails have with sausages, a name given to the plant by the Midwest Polish community who had noticed a striking similarity between the plant and a common Polish dish,
kiełbasa Kielbasa (, ; from Polish ) is any type of meat sausage from Poland and a staple of Polish cuisine. In American English the word typically refers to a coarse, U-shaped smoked sausage of any kind of meat, which closely resembles the ''Wiejska'' ...
.


See also

*
Typha domingensis ''Typha domingensis'', known commonly as southern cattail or cumbungi, is a perennial herbaceous plant of the genus '' Typha''. Distribution and habitat It is found throughout temperate and tropical regions worldwide. It is sometimes found as a ...


References


External links


Can you actually eat cattails?
from ''
The Straight Dope "The Straight Dope" was a question-and-answer newspaper column written under the pseudonym Cecil Adams. Contributions were made by multiple authors, and it was illustrated (also pseudonymously) by Slug Signorino. It was first published in 1973 in ...
'' {{Authority control Aquatic plants Inflorescence vegetables Poales genera Root vegetables