Tympanuchus Cupido
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The greater prairie chicken or pinnated grouse (''Tympanuchus cupido''), sometimes called a boomer,Friederici, Peter (July 20, 1989)
"The Last Prairie Chickens"
''Chicago Reader''. Retrieved August 27, 2014.(Chinese 中文:帕艺明彩大凤凰)
is a large
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
in the
grouse Grouse are a group of birds from the order Galliformes, in the family Phasianidae. Grouse are presently assigned to the tribe Tetraonini (formerly the subfamily Tetraoninae and the family Tetraonidae), a classification supported by mitochondr ...
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
. This
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
n
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
was once abundant, but has become extremely rare and extirpated over much of its range due to
habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
. Conservation measures are underway to ensure the sustainability of existing small populations. One of the most famous aspects of these creatures is the mating ritual called booming.


Description

Adults of both sexes are medium to large chicken-like birds, stocky with round-wings. They have short tails which are typically rounded. Adult males have orange comb-like feathers over their eyes and dark, elongated head feathers that can be raised or lain along neck. They also possess a circular, un-feathered neck patch which can be inflated while displaying; this, like their comb feathers, is also orange. As with many other bird species, the adult females have shorter head feathers and also lack the male's yellow comb and orange neck patch. Adults are about 16.9 in (43 cm) long, and weigh between 24.7–42.3 oz (700–1200 g). The greater prairie-chicken has a wingspan range of 27.4-28.5 in (69.5-72.5 cm).


Subspecies

There are three subspecies; * The
heath hen The heath hen (''Tympanuchus cupido cupido'') is an extinct subspecies of the greater prairie chicken (''Tympanuchus cupido''), a large North American bird in the grouse family. It became extinct in 1932. Heath hens lived in the scrubby heath ...
, ''Tympanuchus cupido cupido,'' which was historically found along the Atlantic coast, is extinct. It was possibly a distinct species; in this case the two other forms would be ''T. pinnatus pinnatus'' and ''T. p. attwateri''. *
Attwater's prairie chicken Attwater's prairie chicken (''Tympanuchus cupido attwateri'') is a highly endangered subspecies of the greater prairie chicken that is native to coastal Texas and formerly Louisiana in the United States. Description Attwater's prairie chicken me ...
, ''T. c. attwateri'' is
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inva ...
and restricted to coastal
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
. * The greater prairie chicken, ''T. c. pinnatus'', is now restricted to a small section of its former range.


Population and habitat

Greater prairie chickens prefer undisturbed
prairie Prairies are ecosystems considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and a composition of grasses, herbs, and shrubs, rather than trees, as the ...
and were originally found in
tallgrass prairie The tallgrass prairie is an ecosystem native to central North America. Historically, natural and anthropogenic fire, as well as grazing by large mammals (primarily bison) provided periodic disturbances to these ecosystems, limiting the encroach ...
s. They can tolerate agricultural land mixed with prairie, but fewer prairie chickens are found in areas that are more agricultural. Their diet consists primarily of seeds and
fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...
, but during the summer they also eat green plants and
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s such as grasshoppers, crickets, and beetles. These birds were once widespread all across the
oak savanna An oak savanna is a type of savanna—or lightly forested grassland—where oaks (''Quercus ''spp.) are the dominant trees. The terms "oakery" or "woodlands" are also used commonly, though the former is more prevalent when referencing the Medite ...
and tall grass prairie
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syste ...
.


Conservation

The greater prairie chicken was almost extinct in the 1930s due to hunting pressure and habitat loss. In
Illinois Illinois ( ) is a U.S. state, state in the Midwestern United States, Midwestern United States. Its largest metropolitan areas include the Chicago metropolitan area, and the Metro East section, of Greater St. Louis. Other smaller metropolita ...
alone, in the 1800s the prairie chicken numbered in the millions. They were a popular game bird, and like many prairie birds, which have also suffered massive habitat loss, they are now on the verge of extinction, with the wild bird population at around 200 in Illinois in 2019. They now only live on small parcels of managed prairie land. Throughout North America, it is thought that their current population has declined severely, to approximately 500,000 individuals. In May 2000, the Canadian Species at Risk Act listed the greater prairie chicken as extirpated in its Canadian range (
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
,
Saskatchewan Saskatchewan ( ; ) is a Provinces and territories of Canada, province in Western Canada, western Canada, bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, to the northeast by Nunavut, and on t ...
,
Manitoba Manitoba ( ) is a Provinces and territories of Canada, province of Canada at the Centre of Canada, longitudinal centre of the country. It is Canada's Population of Canada by province and territory, fifth-most populous province, with a population o ...
,
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...
). It was again confirmed by the Committee on the Status of Endangered Wildlife in Canada in November 2009. Nonetheless, sightings and encounters continue to occur in the south-central regions of Alberta and Saskatchewan, along with southern Ontario, where sightings are extremely rare. In states such as Iowa and Missouri that once had thriving prairie chicken populations (estimated to be hundreds of thousands), total numbers have dropped to about 500. However, the Missouri Department of Conservation has started a program to import prairie chickens from Kansas and Nebraska in the hopes that they will be able to repopulate the state and increase that number to 3,000. Central Wisconsin is home to approximately 600 individuals, down from 55,000 when hunting was prohibited in 1954. Though this area was predominately spruce and tamarack marsh before European settlement, early pioneers drained the marshes and attempted to farm the poor soil. As the prairies to the south and west were lost to agriculture and development, and the southern half of Wisconsin was logged, the prairies spread northward into the abandoned farmland. Today, over 30,000 acres are managed by the Wisconsin Department of Natural Resources as greater prairie chicken habitat. Birdwatchers travel from around the world to visit Wisconsin in April for the Central Wisconsin Prairie Chicken Festival, started in 2006 by Golden Sands Resource Conservation & Development Council, Inc.


Threats

Greater prairie chickens are not threatened by severe winter weather. When the snow is thick they "dive" into the snow to keep warm. A greater threat to the prairie chickens comes in the form of spring rains. These sometimes drenching rains can wreak havoc on their chicks. Another major natural threat is
drought A drought is defined as drier than normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D.  Jiang, A.  Khan, W.  Pokam Mba, D.  Rosenfeld, J. Tierney, an ...
. A drought can destroy food and make it difficult for the chicks. Human interactions are by far the greatest threat. The conversion of native prairie to cropland is very detrimental to these birds. It was found in a radio telemetry study conducted by
Kansas State University Kansas State University (KSU, Kansas State, or K-State) is a public land-grant research university with its main campus in Manhattan, Kansas, United States. It was opened as the state's land-grant college in 1863 and was the first public instit ...
that "most prairie chicken hens avoided nesting or rearing their broods within a quarter-mile of power lines and within a third-mile of improved roads." (Kansas Department of Wildlife and Parks) It was also found that the prairie-chickens avoided communication towers and rural farms. After
population bottleneck A population bottleneck or genetic bottleneck is a sharp reduction in the size of a population due to environmental events such as famines, earthquakes, floods, fires, disease, and droughts; or human activities such as specicide, widespread violen ...
, human management of populations can also produce a loss of
genetic variation Genetic variation is the difference in DNA among individuals or the differences between populations. The multiple sources of genetic variation include mutation and genetic recombination. Mutations are the ultimate sources of genetic variation, ...
and
genetic diversity Genetic diversity is the total number of genetic characteristics in the genetic makeup of a species, it ranges widely from the number of species to differences within species and can be attributed to the span of survival for a species. It is dis ...
in the species.


Sexual behavior

Greater prairie chickens do not migrate. They are territorial birds and often defend their booming grounds. These booming grounds are the area in which they perform their displays in hopes of attracting females. Their displays consist of inflating air sacs located on the side of their neck and snapping their tails. These booming grounds usually have very short or no vegetation. The male prairie-chickens stay on this ground displaying for almost two months. The breeding season usually begins in the United States starting in late March throughout April. During this time the males establish booming sites where they display for the females. The one or two most dominant males can obtain 90% of mating opportunities. Due to their now small populations and
habitat fragmentation Habitat fragmentation describes the emergence of discontinuities (fragmentation) in an organism's preferred environment (habitat), causing population fragmentation and ecosystem decay. Causes of habitat fragmentation include geological processes ...
the greater prairie chickens often undergo inbreeding causing observable
inbreeding depression Inbreeding depression is the reduced biological fitness which has the potential to result from inbreeding (the breeding of related individuals). Biological fitness refers to an organism's ability to survive and perpetuate its genetic material. In ...
: with fewer offspring and a decreased survival rate within these limited offspring further aiding their population decrease. After mating has taken place, the females move about one mile from the booming grounds and begin to build their
nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materia ...
s. Hens lay between 5 and 17
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...
s per clutch and the eggs take between 23 and 24 days to hatch. There are between five and 10 young per brood. (INRIN, 2005). The young are raised by the female and
fledge Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnerable c ...
in one to four weeks, are completely independent by the tenth to twelfth week, and reach sexual maturity by age one (Ammann, 1957). A study of female greater prairie chickens in Kansas found that their survival rates were 1.6 to 2.0 times higher during the non-breeding season compared to the breeding season; this was due to heavy predation during nesting and brood-rearing.Augustine JK, Sandercock BK (2011) ''Demography of female Greater Prairie-Chickens in unfragmented grasslands in Kansas''. Avian Conservation and Ecology 6(1):2

One problem facing prairie chickens is competition with the
ring-necked pheasant The common pheasant (''Phasianus colchicus'') is a bird in the pheasant family (Phasianidae). The genus name comes from Latin ''phasianus'', "pheasant". The species name ''colchicus'' is Latin for "of Colchis" (modern day Georgia), a country on ...
s. Pheasants lay their eggs in prairie-chicken nests. The pheasant eggs hatch first; this causes the prairie chickens to leave the nest thinking that the young have hatched. In reality the eggs did not hatch and the young usually die because the mother is not there to incubate the eggs. File:Greater Prairie Chicken -male displaying and spinning.theora.ogv File:Greater Prairie Chicken - males displaying to a female 700.theora.ogv File:Tympanuchus cupido -Illinois, USA -male jumps and displays-8.ogv


See also

*
Lesser prairie chicken The lesser prairie chicken (''Tympanuchus pallidicinctus'') is a species in the grouse family. Description It is a medium to large bird, striped white and brown, slightly smaller and paler than its near relative the greater prairie chicken (''T. ...
* Lekking


References

* ARKive
images and movies of the greater prairie chicken ''(Tympanuchus cupido)''
* USGS Patuxent Bird Identification InfoCenter

* Gunderson, Dan.
Prairie chickens booming again.
Minnesota Public Radio (2006) * Ammann, G. A. 1957 The prairie grouse of Michigan. Michigan Dept. Consew. Tech. Bull. * Illinois Natural Resource Information Network

* Kansas Department of Wildlife and Parks
Greater and Lesser Prairie Chickens

State conservationists scour the Kansas boondocks, aiming to repopulate Missouri with horny prairie chickens


External links


Comparative Analysis between the Greater Prairie Chicken and the Extinct Heath Hen




* ttps://web.archive.org/web/20110121202756/http://www.nature.org/animals/birds/animals/gprairiechicken.html The Nature Conservancy's Grassland Birds: Greater Prairie Chicken
gbwf.org - Greater Prairie Chicken

eNature.com - Greater Prairie Chicken


(for
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...
)
Greater Prairie Chicken photo gallery
VIREO
The Return of the Prairie Chicken
Documentary produced by
Iowa Public Television Iowa PBS, formerly Iowa Public Television (IPTV), is a network of Public Broadcasting Service (PBS) member stations in the U.S. state of Iowa. It is owned by the Iowa Public Broadcasting Board, an agency of the state education department which h ...
{{Taxonbar, from=Q19630 Tympanuchus Grouse Endemic birds of the Plains-Midwest (United States) Birds described in 1758 Articles containing video clips Taxa named by Carl Linnaeus