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In the
anatomy Anatomy () is the branch of biology concerned with the study of the structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having its ...
of humans and various other
tetrapod Tetrapods (; ) are four-limb (anatomy), limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant taxon, extant and extinct amphibians, sauropsids (reptiles, including dinosaurs and therefore birds) and synapsids (p ...
s, the eardrum, also called the tympanic membrane or myringa, is a thin, cone-shaped
membrane A membrane is a selective barrier; it allows some things to pass through but stops others. Such things may be molecules, ions, or other small particles. Membranes can be generally classified into synthetic membranes and biological membranes. ...
that separates the external ear from the
middle ear The middle ear is the portion of the ear medial to the eardrum, and distal to the oval window of the cochlea (of the inner ear). The mammalian middle ear contains three ossicles, which transfer the vibrations of the eardrum into waves in ...
. Its function is to transmit
sound In physics, sound is a vibration that propagates as an acoustic wave, through a transmission medium such as a gas, liquid or solid. In human physiology and psychology, sound is the ''reception'' of such waves and their ''perception'' by ...
from the air to the ossicles inside the middle ear, and then to the oval window in the fluid-filled
cochlea The cochlea is the part of the inner ear involved in hearing. It is a spiral-shaped cavity in the bony labyrinth, in humans making 2.75 turns around its axis, the modiolus. A core component of the cochlea is the Organ of Corti, the sensory org ...
. Hence, it ultimately converts and amplifies vibration in the air to vibration in cochlear fluid. The
malleus The malleus, or hammer, is a hammer-shaped small bone or ossicle of the middle ear. It connects with the incus, and is attached to the inner surface of the eardrum. The word is Latin for 'hammer' or 'mallet'. It transmits the sound vibrations ...
bone bridges the gap between the eardrum and the other ossicles. Rupture or perforation of the eardrum can lead to conductive hearing loss. Collapse or
retraction Retraction or retract(ed) may refer to: Academia * Retraction in academic publishing, withdrawals of previously published academic journal articles Mathematics * Retraction (category theory) * Retract (group theory) * Retraction (topology) Huma ...
of the eardrum can cause conductive hearing loss or cholesteatoma.


Structure


Orientation and relations

The tympanic membrane is oriented obliquely in the anteroposterior, mediolateral, and superoinferior planes. Consequently, its superoposterior end lies lateral to its anteroinferior end. Anatomically, it relates superiorly to the middle cranial fossa, posteriorly to the ossicles and
facial nerve The facial nerve, also known as the seventh cranial nerve, cranial nerve VII, or simply CN VII, is a cranial nerve that emerges from the pons of the brainstem, controls the muscles of facial expression, and functions in the conveyance of ta ...
, inferiorly to the
parotid gland The parotid gland is a major salivary gland in many animals. In humans, the two parotid glands are present on either side of the mouth and in front of both ears. They are the largest of the salivary glands. Each parotid is wrapped around the m ...
, and anteriorly to the temporomandibular joint.


Regions

The eardrum is divided into two general regions: the pars flaccida and the pars tensa. The relatively fragile pars flaccida lies above the lateral process of the
malleus The malleus, or hammer, is a hammer-shaped small bone or ossicle of the middle ear. It connects with the incus, and is attached to the inner surface of the eardrum. The word is Latin for 'hammer' or 'mallet'. It transmits the sound vibrations ...
between the notch of Rivinus and the anterior and posterior malleal folds. Consisting of two layers and appearing slightly pinkish in hue, it is associated with
Eustachian tube In anatomy, the Eustachian tube, also known as the auditory tube or pharyngotympanic tube, is a tube that links the nasopharynx to the middle ear, of which it is also a part. In adult humans, the Eustachian tube is approximately long and in ...
dysfunction and cholesteatomas. The larger pars tensa consists of three layers:
skin Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation. Other cuticle, animal coverings, such as the arthropod exoskeleton, have diffe ...
,
fibrous tissue Fiber or fibre (from la, fibra, links=no) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorpora ...
, and mucosa. Its thick periphery forms a fibrocartilaginous ring called the annulus tympanicus or Gerlach's ligament. while the central umbo tents inward at the level of the tip of malleus. The middle fibrous layer, containing radial, circular, and parabolic fibers, encloses the handle of malleus. Though comparatively robust, the pars tensa is the region more commonly associated with perforations.


Umbo

The manubrium ( la, handle) of the
malleus The malleus, or hammer, is a hammer-shaped small bone or ossicle of the middle ear. It connects with the incus, and is attached to the inner surface of the eardrum. The word is Latin for 'hammer' or 'mallet'. It transmits the sound vibrations ...
is firmly attached to the medial surface of the membrane as far as its center, drawing it toward the tympanic cavity. The lateral surface of the membrane is thus concave. The most depressed aspect of this concavity is termed the umbo ( la, shield boss).


Nerve supply

Sensation of the outer surface of the tympanic membrane is supplied mainly by the
auriculotemporal nerve The auriculotemporal nerve is a branch of the mandibular nerve (CN V3) that runs with the superficial temporal artery and vein, and provides sensory innervation to various regions on the side of the head. Structure Origin The auriculotemp ...
, a branch of the mandibular nerve (
cranial nerve Cranial nerves are the nerves that emerge directly from the brain (including the brainstem), of which there are conventionally considered twelve pairs. Cranial nerves relay information between the brain and parts of the body, primarily to and f ...
V3), with contributions from the auricular branch of the vagus nerve ( cranial nerve X), the
facial nerve The facial nerve, also known as the seventh cranial nerve, cranial nerve VII, or simply CN VII, is a cranial nerve that emerges from the pons of the brainstem, controls the muscles of facial expression, and functions in the conveyance of ta ...
(cranial nerve VII), and possibly the glossopharyngeal nerve (cranial nerve IX). The inner surface of the tympanic membrane is innervated by the glossopharyngeal nerve.


Clinical significance


Examination

When the eardrum is illuminated during a medical examination, a cone of light radiates from the tip of the malleus to the periphery in the anteroinferior quadrant, this is what is known clinically as 5 o'clock.


Rupture

Unintentional perforation (rupture) has been described in
blast injuries A blast injury is a complex type of physical trauma resulting from direct or indirect exposure to an explosion. Blast injuries occur with the detonation of high-order explosives as well as the deflagration of low order explosives. These inju ...
and air travel, typically in patients experiencing upper respiratory congestion that prevents equalization of pressure in the middle ear. It is also known to occur in swimming, diving (including
scuba diving Scuba diving is a mode of underwater diving whereby divers use breathing equipment that is completely independent of a surface air supply. The name "scuba", an acronym for "Self-Contained Underwater Breathing Apparatus", was coined by Chri ...
), and martial arts. Patients with tympanic membrane rupture may experience bleeding,
tinnitus Tinnitus is the perception of sound when no corresponding external sound is present. Nearly everyone experiences a faint "normal tinnitus" in a completely quiet room; but it is of concern only if it is bothersome, interferes with normal hearin ...
,
hearing loss Hearing loss is a partial or total inability to hear. Hearing loss may be present at birth or acquired at any time afterwards. Hearing loss may occur in one or both ears. In children, hearing problems can affect the ability to acquire spoken la ...
, or disequilibrium (
vertigo Vertigo is a condition where a person has the sensation of movement or of surrounding objects moving when they are not. Often it feels like a spinning or swaying movement. This may be associated with nausea, vomiting, sweating, or difficulties ...
). However, they rarely require medical intervention, as between 80 and 95 percent of ruptures recover completely within two to four weeks. The prognosis becomes more guarded as the force of injury increases.


Surgical puncture for treatment of middle ear infections

The pressure of fluid in an infected middle ear onto the eardrum may cause it to rupture. Usually, this consists of a small hole (perforation), which allows fluid to drain out. If this does not occur naturally, a myringotomy (tympanotomy, tympanostomy) can be performed. A myringotomy is a surgical procedure in which a tiny incision is created in the eardrum to relieve pressure caused by excessive buildup of fluid, or to drain pus from the
middle ear The middle ear is the portion of the ear medial to the eardrum, and distal to the oval window of the cochlea (of the inner ear). The mammalian middle ear contains three ossicles, which transfer the vibrations of the eardrum into waves in ...
. The fluid or pus comes from a middle ear infection (
otitis media Otitis media is a group of inflammatory diseases of the middle ear. One of the two main types is acute otitis media (AOM), an infection of rapid onset that usually presents with ear pain. In young children this may result in pulling at the ear, ...
), which is a common problem in children. A
tympanostomy tube Tympanostomy tube, also known as a grommet or myringotomy tube, is a small tube inserted into the eardrum in order to keep the middle ear aerated for a prolonged period of time, and to prevent the accumulation of fluid in the middle ear. The ope ...
is inserted into the eardrum to keep the middle ear aerated for a prolonged time and to prevent reaccumulation of fluid. Without the insertion of a tube, the incision usually heals spontaneously in two to three weeks. Depending on the type, the tube is either naturally extruded in 6 to 12 months or removed during a minor procedure. Those requiring myringotomy usually have an obstructed or dysfunctional
eustachian tube In anatomy, the Eustachian tube, also known as the auditory tube or pharyngotympanic tube, is a tube that links the nasopharynx to the middle ear, of which it is also a part. In adult humans, the Eustachian tube is approximately long and in ...
that is unable to perform drainage or ventilation in its usual fashion. Before the invention of antibiotics, myringotomy without tube placement was also used as a major treatment of severe acute otitis media. In some cases, the pressure of fluid in an infected middle ear is great enough to cause the eardrum to rupture naturally. Usually, this consists of a small hole (perforation), from which fluid can drain.


Society and culture

The Bajau people of the
Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
intentionally rupture their eardrums at an early age to facilitate diving and hunting at sea. Many older Bajau therefore have difficulties hearing.


See also

*
Middle ear The middle ear is the portion of the ear medial to the eardrum, and distal to the oval window of the cochlea (of the inner ear). The mammalian middle ear contains three ossicles, which transfer the vibrations of the eardrum into waves in ...
* Valsalva maneuver to equalize pressure across the eardrum


Additional images

File:Anatomy of the Human Ear en.svg, Anatomy of the human right ear. File:Gray907.png, External and middle ear, right side, opened from the front (coronal section) File:Gray908.png, Horizontal section through left ear; upper half of section File:Gray912.png, The right membrana tympani with the hammer and the chorda tympani, viewed from within, from behind, and from above File:Gray915.png, Auditory tube, laid open by a cut in its long axis File:Gray919.png, Chain of ossicles and their ligaments, seen from the front in a vertical, transverse section of the tympanum ympanic cavityFile:Gray909.png, Right eardrum as seen through a speculum File:Normal Left Tympanic Membrane.jpg, This is a normal left eardrum. File:Tympanic membrane viewed by otoscope.png, Tympanic membrane viewed by otoscope File:Traumatic Perforation of the Tympanic Membrane.jpg, The oval perforation in this left tympanic membrane was the result of a slap on the ear File:Subtotal Perforation of the right tympanic membrane.tif, A subtotal perforation of the right tympanic membrane resulting from a previous severe otitis media File:TM RIGHT NORMAL.jpg, A normal human right tympanic membrane (eardrum) File:Frog on leaf with eardrum.jpg, Frog on leaf showing eardrum


References


External links


Diagram at Georgia State University


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