Turnstones are two

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

that comprise the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

''Arenaria'' in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Scolopacidae Sandpipers are a large family, Scolopacidae, of waders. They include many species called sandpipers, as well as those called by names such as curlew and snipe. The majority of these species eat small invertebrates picked out of the mud or soil. ...

. They are closely related to

calidrid ''Calidris'' is a genus of Arctic-breeding, strongly migratory wading birds in the family Scolopacidae. These birds form huge mixed flocks on coasts and estuaries in winter. They are the typical "sandpipers", small to medium-sized, long-winged ...

sandpipers and might be considered members of the

tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English language, English is in the discipline of anthropology. This definition is contested, in p ...

Calidriini. The

''Arenaria'' was introduced by the French zoologist

Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published works ...

in 1760 with the

ruddy turnstone The ruddy turnstone (''Arenaria interpres'') is a small cosmopolitan wading bird, one of two species of turnstone in the genus ''Arenaria''. It is now classified in the sandpiper family Scolopacidae but was formerly sometimes placed in the plov ...

(''Arenaria interpres'') as the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

. The genus name ''arenaria'' is from

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

''arenarius'', "inhabiting sand", from ''arena'', "sand". The genus contains two species: the

(''Arenaria interpres'') and the

black turnstone The black turnstone (''Arenaria melanocephala'') is a species of small wading bird. It is one of two species of turnstone in the genus ''Arenaria'' the ruddy turnstone (''A. interpres'') being the other. It is now classified in the sandpiper fa ...

(''Arenaria melanocephala''). Both birds are

wader 245px, A flock of Dunlins and Red knots">Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflat ...

s. Their length is typically between 20 and 25 cm, with a wingspan between 50 and 60 cm and a body mass between 110 and 130g. For waders their build is stocky, with short, slightly upturned, wedge shaped bills. They have white patches on the back, wings and tail. They are high

Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...

breeders, and are migratory. Their strong necks and powerful, slightly upturned bills are adapted to their feeding technique. As the name implies, these species overturn stones, seaweed, and similar items in search of

invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...

prey.Svensson, Lars et al. Collins Bird Guide 2nd ed. Publisher: Collins 2010. They are strictly coastal, prefer stony beaches to sand, and often share beach space with other species of waders such as

purple sandpiper The purple sandpiper (''Calidris maritima'') is a small shorebird in the sandpiper family Scolopacidae. This is a hardy sandpiper that breeds in the arctic and subarctic regions of Eurasia and North America and winters further south on the Atlant ...

Species

Ruddy turnstone

The

(or just turnstone in

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...

), ''Arenaria interpres'', has a circumpolar distribution, and is a very long distance migrant, wintering on coasts as far south as

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

and

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

. It is thus a common sight on coasts almost everywhere in the world. It is one of the species to which the ''Agreement on the Conservation of African-Eurasian Migratory Waterbirds'' (

AEWA The Agreement on the Conservation of African-Eurasian Migratory Waterbirds, or African-Eurasian Waterbird Agreement (AEWA) is an independent international treaty developed under the auspices of the United Nations Environment Programme's Conventio ...

) applies. In breeding

plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...

, this is a showy bird, with a black-and-white head, chestnut back, white underparts and red legs. The drabber winter plumage is basically brown above and white below. This is a generally tame bird and is an opportunist feeder. Unlike most waders, it will scavenge, and has a phenomenal list of recorded food items, including human corpses and

coconut The coconut tree (''Cocos nucifera'') is a member of the palm tree family ( Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, the seed, or the ...

. The call is a staccato ''tuck- tuck- tuck''.

Black turnstone

The

(''Arenaria melanocephala'') has a similar structure to its widespread relative, but has black upperparts and chest, and white below. It has a much more restricted range than the ruddy turnstone, breeding in western

Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...

, and wintering mainly on the

Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...

coast of the

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

. There exists a

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

bone, a

distal Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...

piece of tarsometatarsus found in the

Edson Beds Edson may refer to: Places Canada * Edson, Alberta United States * Edson, Kansas, an unincorporated community * Edson, South Dakota, a ghost town * Edson, Wisconsin, a town ** Edson (community), Wisconsin, an unincorporated community People ...

Sherman County, Kansas Sherman County (standard abbreviation: SH) is a county located in the U.S. state of Kansas. As of the 2020 census, the county population was 5,927. Its county seat is Goodland. Sherman County was created by the Legislature of 1873, and named ...

. Dating from the mid- Blancan some 4-3 million years ago, it appears to be from a calidriid somewhat similar to a

pectoral sandpiper The pectoral sandpiper (''Calidris melanotos'') is a small, migratory wader that breeds in North America and Asia, wintering in South America and Oceania. It eats small invertebrates. Its nest, a hole scraped in the ground and with a thick linin ...

, but has some traits reminiscent of turnstones. Depending on which traits are

apomorph In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have ...

ic and

plesiomorph In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, a ...

ic, it may be an ancestral representative of either lineage.

References

External links

Ruddy turnstone
– Cornell Lab of Ornithology

– USGS Patuxent Bird Identification InfoCenter
Turnstones feeding and bathing
{{Taxonbar, from=Q4865776 Sandpipers Taxa named by Mathurin Jacques Brisson