''Tuerkayana hirtipes'' is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of terrestrial crab.

Description and behaviour

''T. hirtipes'' can be distinguished from members of the genus ''

Discoplax ''Discoplax'' is a genus of terrestrial crabs. It is very closely related to the genus ''Cardisoma''. Five species are currently recognised: *''Discoplax celeste'' 2012 *'' Discoplax gracilipes'' Ng & Guinot, 2001 *'' Discoplax longipes'' A. Mi ...

'' by the smooth and distinctly inflated

carapace A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tor ...

. It is also blue or blue-brown in colour, compared to purple or purple-brown in the remaining species. The breeding season for ''T. hirtipes'' lasts seven months, and females must migrate to the sea to release their

larvae A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...

Distribution

''T. hirtipes'' has a wide distribution across the western

Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...

. It has been recorded from southern

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...

Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...

, southern

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Palau Palau,, officially the Republic of Palau and historically ''Belau'', ''Palaos'' or ''Pelew'', is an island country and microstate in the western Pacific. The nation has approximately 340 islands and connects the western chain of the Caro ...

Guam Guam (; ch, Guåhan ) is an organized, unincorporated territory of the United States in the Micronesia subregion of the western Pacific Ocean. It is the westernmost point and territory of the United States (reckoned from the geographic cent ...

, eastern

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

Fiji Fiji ( , ,; fj, Viti, ; Fiji Hindi: फ़िजी, ''Fijī''), officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists ...

Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...

(a single record, almost certainly erroneous), the

Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capita ...

New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...

, the

Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...

Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...

Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...

Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...

and

Singapore Singapore (), officially the Republic of Singapore, is a sovereign island country and city-state in maritime Southeast Asia. It lies about one degree of latitude () north of the equator, off the southern tip of the Malay Peninsula, borde ...

. Records from the Indian Ocean are now considered to belong to two related species, '' T. celeste'' ( Christmas Island only) and '' D. magna'' (relatively widespread in eastern Indian Ocean).

Taxonomic history

''Tuerkayana hirtipes'' was originally described by James Dwight Dana under the name ''Cardisoma hirtipes''. His description was published in 1851 based on material collected on the

United States Exploring Expedition The United States Exploring Expedition of 1838–1842 was an exploring and surveying expedition of the Pacific Ocean and surrounding lands conducted by the United States. The original appointed commanding officer was Commodore Thomas ap Catesby ...

("Wilkes expedition") in Fiji. The

type material In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the ...

has been lost, so a neotype from Fiji was designated in 2012. In late 2018, a re-examination of the taxonomy of

Gecarcinidae The Gecarcinidae, the land crabs, are a family of true crabs that are adapted for terrestrial existence. Similar to all other crabs, land crabs possess a series of gills. In addition, the part of the carapace covering the gills is inflated and eq ...

resulted in the creation of the genus ''Tuerkayana'', which was created with four species from ''

Cardisoma ''Cardisoma'' is a genus of large land crabs. Three species formerly placed in this genus are now placed in '' Discoplax''. The four species that remain in ''Cardisoma'' are found in warm coastal regions where they live in burrows. Young individu ...

'' and ''

'', including ''hirtipes''.

References

{{Taxonbar, from=Q5281680 Grapsoidea Terrestrial crustaceans Crustaceans described in 1851