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Tsintaosaurini is a tribe of basal
lambeosaurine Lambeosaurinae is a group of crested hadrosaurid dinosaurs. Classification Lambeosaurines have been traditionally split into the tribes or clades Parasaurolophini ('' Parasaurolophus'', ''Charonosaurus'', others (?).) and Lambeosaurini (''Cor ...
hadrosaurs Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which incl ...
native to
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
. It is thought to contains the genera ''
Tsintaosaurus ''Tsintaosaurus'' (; meaning "Qingdao lizard", after the old transliteration "Tsingtao") is a genus of hadrosaurid dinosaur from China. It was about long and weighed . The type species is ''Tsintaosaurus spinorhinus'', first described by Chine ...
'' (from
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
), ''
Pararhabdodon ''Pararhabdodon'' (meaning "near fluted tooth" in reference to ''Rhabdodon'') is a genus of tsintaosaurin hadrosaurid dinosaur, from the Maastrichtian-age Upper Cretaceous Tremp Group of Spain. The first remains were discovered from the Sant R ...
'' (from
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
) and '' Koutalisaurus'' (also from Spain), though some studies have questioned its existence as a natural grouping.


Classification

Tsintaosaurini is part of the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Hadrosauridae, specifically the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Lambeosaurinae Lambeosaurinae is a group of crested hadrosaurid dinosaurs. Classification Lambeosaurines have been traditionally split into the tribes or clades Parasaurolophini ('' Parasaurolophus'', '' Charonosaurus'', others (?).) and Lambeosaurini ('' C ...
. The existence of a tsintaosaur clade of lambeosaurines was first recognized by palaeontologists Albert Prieto-Márquez and Johnathan R. Wagner, who in 2009 published a paper recognizing a
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
relationship between ''
Tsintaosaurus ''Tsintaosaurus'' (; meaning "Qingdao lizard", after the old transliteration "Tsingtao") is a genus of hadrosaurid dinosaur from China. It was about long and weighed . The type species is ''Tsintaosaurus spinorhinus'', first described by Chine ...
'' and ''
Pararhabdodon ''Pararhabdodon'' (meaning "near fluted tooth" in reference to ''Rhabdodon'') is a genus of tsintaosaurin hadrosaurid dinosaur, from the Maastrichtian-age Upper Cretaceous Tremp Group of Spain. The first remains were discovered from the Sant R ...
'' based both on shared anatomical traits and a
phylogenetic analysis In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
. A 2013 study by Prieto-Márquez corroborated the existence of this grouping, and coined the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English language, English is in the discipline of anthropology. This definition is contested, in p ...
Tsintaosaurini to refer to it. The
type genus In biological taxonomy, the type genus is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearing type of a nominal f ...
is ''Tsintaosaurus'', and it was defined as the smallest
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
containing ''Tsintaosaurus spinorhinus'' and ''Pararhabdodon isonensis''. Several studies since have corroborated the existence of the clade, though some others have failed to recover it, instead finding the two genera in a
polytomy An internal node of a phylogenetic tree is described as a polytomy or multifurcation if (i) it is in a rooted tree and is linked to three or more child subtrees or (ii) it is in an unrooted tree and is attached to four or more branches. A tr ...
of basal lambeosaurs. A 2021 paper by Daniel Madzia and other ornithischian researchers, focused on a revising ornithischian nomenclature and converting existing group names into
Phylocode The ''International Code of Phylogenetic Nomenclature'', known as the ''PhyloCode'' for short, is a formal set of rules governing phylogenetic nomenclature. Its current version is specifically designed to regulate the naming of clades, leaving the ...
-compliant clades, re-formalized the coining of Tsintaosaurini and revised its definition to be the most inclusive group including ''T. spinorhinus'' and ''P. isonensis'', but not '' Aralosaurus tuberiferus'', ''
Lambeosaurus lambei ''Lambeosaurus'' ( , meaning " Lambe's lizard") is a genus of hadrosaurid dinosaur that lived about 75 million years ago, in the Late Cretaceous period (Campanian stage) of North America. This bipedal/quadrupedal, herbivorous dinosaur is kn ...
'', or ''
Parasaurolophus walkeri ''Parasaurolophus'' (; meaning "near crested lizard" in reference to ''Saurolophus)'' is a genus of herbivorous hadrosaurid ornithopod dinosaur that lived in what is now North America and possibly Asia during the Late Cretaceous Period, about ...
''. The following cladogram shows the results of a phylogenetic analysis of hadrosaur relationships from a 2022 study: A 2020 study by Nick Longrich and colleagues describing the genus '' Ajnabia'' found ''Pararhabdodon'' to be part a
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
clade of European lambeosaurs, termed Arenysaurini, rather than a relative of ''Tsintaosaurus'', thereby making the tribe an inapplicable
polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of converg ...
grouping. The cladogram from their study's phylogenetic analysis is shown below:


See also

*
Timeline of hadrosaur research A timeline is a display of a list of events in chronological order. It is typically a graphic design showing a long bar labelled with dates paralleling it, and usually contemporaneous events. Timelines can use any suitable scale represent ...


References

Lambeosaurines {{ornithopod-stub