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True gobies were a
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
, the Gobiinae, of the goby
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Gobiidae Gobiidae or gobies is a family of bony fish in the order Gobiiformes, one of the largest fish families comprising more than 2,000 species in more than 200 genera. Most of gobiid fish are relatively small, typically less than in length, and the ...
, although the 5th edition of the ''
Fishes of the World ''Fishes of the World'' by the American ichthyologist Joseph S. Nelson (1937–2011) is a standard reference for fish systematics. Now in its fifth edition (2016), the work is a comprehensive overview of the diversity and classification of the ...
'' does not subdivide the Gobiidae into subfamilies. They are found in all
ocean The ocean (also the sea or the world ocean) is the body of salt water that covers approximately 70.8% of the surface of Earth and contains 97% of Earth's water. An ocean can also refer to any of the large bodies of water into which the wo ...
s and a few
river A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, sea, lake or another river. In some cases, a river flows into the ground and becomes dry at the end of its course without reaching another body of wate ...
s and
lake A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much large ...
s, but most live in warm waters. Altogether, the Gobiinae unite about 1149 described
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
in 160
genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
, and new ones are still being discovered in numbers.


Description and ecology

They are usually mid-sized to small
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...
es; some are very colorful, while others are
cryptic Cryptic may refer to: In science: * Cryptic species complex, a group of species that are very difficult to distinguish from one another * Crypsis, the ability of animals to blend in to avoid observation * Cryptic era, earliest period of the Earth ...
. Most true gobies are less than 10 cm (4 in) long when fully grown. The largest species ''
Glossogobius giuris ''Glossogobius giuris'', the tank goby, is a species of goby native to fresh, marine and brackish waters from the Red Sea and East Africa through South Asia and the Indian Ocean to China, Australia and the islands of the Pacific Ocean. This spe ...
'' can reach up to 50 cm (20 in); the smallest known species as of 2010, '' Trimmatom nanus'', is just about 1 cm in length when fully grown, making it one of the smallest
vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...
s. In many true gobies, the
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two en ...
s have grown together into a suction cup they can use to hold on to substrate. Most have two
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
s, the first made up from spiny
fin ray Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as see ...
s, while the other has some spines in the front followed by numerous soft rays. They are most plentiful in the
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
and
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
regions, but as a group are almost
cosmopolitan Cosmopolitan may refer to: Food and drink * Cosmopolitan (cocktail), also known as a "Cosmo" History * Rootless cosmopolitan, a Soviet derogatory epithet during Joseph Stalin's anti-Semitic campaign of 1949–1953 Hotels and resorts * Cosmopoli ...
in
marine ecosystem Marine ecosystems are the largest of Earth's aquatic ecosystems and exist in waters that have a high salt content. These systems contrast with freshwater ecosystems, which have a lower salt content. Marine waters cover more than 70% of the surf ...
s. A few
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
tolerate
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
water, and some – ''
Padogobius ''Padogobius'' is a genus of fish in the family Gobiidae, the gobies. They are native to fresh waters of southern Europe. Species There are currently two recognized species in this genus: * ''Padogobius bonelli'' ( Bonaparte, 1846) (Padanian gob ...
'' and ''
Pomatoschistus ''Pomatoschistus'' is a genus of gobies native to fresh, brackish and marine waters of Europe, the eastern Atlantic Ocean and the Mediterranean Sea. Species There are currently 13 recognized species in this genus: * '' Pomatoschistus anatoliae' ...
'' species – even inhabit
fresh water Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
. They are generally
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...
as adults (the
spawn Spawn or spawning may refer to: * Spawn (biology), the eggs and sperm of aquatic animals Arts, entertainment, and media * Spawn (character), a fictional character in the comic series of the same name and in the associated franchise ** '' Spawn: ...
can distribute widely by
ocean current An ocean current is a continuous, directed movement of sea water generated by a number of forces acting upon the water, including wind, the Coriolis effect, breaking waves, cabbeling, and temperature and salinity differences. Depth contours, s ...
s), only ''Sufflogobius bibarbatus'' is noted to be quite
pelagic The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or wa ...
throughout its life. Most inhabit some sort of burrow or crevice and are somewhat
territorial A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
. In some cases, they live in
symbioses Symbiosis (from Greek , , "living together", from , , "together", and , bíōsis, "living") is any type of a close and long-term biological interaction between two different biological organisms, be it mutualistic, commensalistic, or parasit ...
with unrelated animals, such as
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s. The larger species are fished for food, in some cases on a commercial scale. Many Gobiinae species are popular
aquarium An aquarium (plural: ''aquariums'' or ''aquaria'') is a vivarium of any size having at least one transparent side in which aquatic plants or animals are kept and displayed. Fishkeepers use aquaria to keep fish, invertebrates, amphibians, aq ...
fish. Especially popular are the colorful species, some of which are regularly traded. In general, the interesting behavior and bold habits make most true gobies seem attractive pets. However, their territoriality and because even the smallest species are fundamentally
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
and need living food to thrive make them not easy to keep (particularly compared to the related family
Eleotridae Eleotridae is a family of fish commonly known as sleeper gobies, with about 34 genera and 180 species. Most species are found in the tropical Indo-Pacific region, but there are also species in subtropical and temperate regions, warmer parts of th ...
). As typical for oceanic fishes, many Gobiinae tend to be almost impossible to breed in captivity, and some species have become rare from
habitat destruction Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
and
overfishing Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing fish stock), resulting in th ...
.


Genera

This subfamily contains about 160 genera and 1120 species: * '' Aboma'' * ''
Acentrogobius ''Acentrogobius'' is a genus of gobies native to marine, fresh and brackish waters of the coasts of the Indian Ocean and the western Pacific Ocean. ''Acentrogobius matsya'' is an otolith-based fossil species found in the Burdigalian (Miocene) Qu ...
'' * ''
Afurcagobius ''Afurcagobius'' is a small genus of gobies endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a ...
'' * ''
Akko Acre ( ), known locally as Akko ( he, עַכּוֹ, ''ʻAkō'') or Akka ( ar, عكّا, ''ʻAkkā''), is a city in the coastal plain region of the Northern District of Israel. The city occupies an important location, sitting in a natural harb ...
'' * ''
Amblyeleotris ''Amblyeleotris'' is a genus of fish in the family Gobiidae found throughout the Indo-Pacific region. This is the largest genus of the shrimp gobies or prawn gobies, so-called because of their symbiotic relationship with certain alpheid shrimps. ...
'' * '' Amblygobius'' * '' Amoya'' * ''
Anatirostrum The duckbill pugolovka (''Anatirostrum profundorum'') is a species of gobiid fish endemic to the southern part of the Caspian Sea. It is characterized by its special flattened and elongated head that resembles a duck's bill.Brian W. CoadFreshwat ...
'' * '' Ancistrogobius'' * '' Antilligobius'' * '' Aphia'' * '' Arcygobius'' * '' Arenigobius'' * '' Aruma'' * '' Asterropteryx'' * '' Aulopareia'' * '' Austrolethops'' * ''
Babka A babka is a sweet braided bread (not a cake) which originated in the Jewish communities of Poland and Ukraine. It is popular in Israel (often referred to as simply a yeast cake: ) and in the Jewish diaspora. It is prepared with a yeast-leavened ...
'' * '' Barbulifer'' * '' Barbuligobius'' * ''
Bathygobius ''Bathygobius'' is a circumtropical genus of fish in the family Gobiidae. Abstract Species There are currently 29 recognized species in this genus: * '' Bathygobius aeolosoma'' ( J. D. Ogilby, 1889) * '' Bathygobius albopunctatus'' (Valencienne ...
'' * ''
Benthophiloides ''Benthophiloides'' is a genus of gobies widespread in the basins of the Black Sea and the Caspian Sea The Caspian Sea is the world's largest inland body of water, often described as the world's largest lake or a full-fledged sea. An endorhe ...
'' * ''
Benthophilus The tadpole-gobies (''Benthophilus''), also called pugolovkas (which means "tadpole" in Ukrainian), are a genus of Ponto-Caspian fishes in the family Gobiidae. They are distributed in the fresh and brackish waters of basins of the Black Sea, C ...
'' * '' Bollmannia'' * '' Bryaninops'' * '' Buenia'' * '' Cabillus'' * '' Caffrogobius'' * '' Callogobius'' * ''
Caspiosoma ''Caspiosoma caspium'' is a species of Ponto-Caspian goby native to the deltas of rivers inflows to the north-western Black Sea: Dnieper up to Berislav, also in the Dnieper-Bug Estuary and Berezan Estuary, Danube, Dniester with the estuary, Cuci ...
'' * '' Chriolepis'' * '' Chromogobius'' * '' Corcyrogobius'' * '' Coryogalops'' * ''
Coryphopterus ''Coryphopterus'' is a genus of gobies primarily found in the western Atlantic Ocean, although some species are found in the Indian and/or Pacific oceans. Species There are currently 14 recognized species in this genus: * '' Coryphopterus alloid ...
'' * '' Cristatogobius'' * '' Croilia'' * ''
Cryptocentroides ''Cryptocentroides'' is a genus of gobies native to the western Indian Ocean and the western Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north ...
'' * '' Cryptocentrus'' * '' Crystallogobius'' * '' Cryptopsilotris'' * '' Ctenogobiops'' * '' Deltentosteus'' * '' Didogobius'' * '' Discordipinna'' * '' Dotsugobius'' Shibukawa, K., Suzuki, T. & Senou, H. (2014)
''Dotsugobius'', a New Genus for ''Lophogobius bleekeri'' Popta, 1921 (Actinopterygii, Gobioidei, Gobiidae), with Re-description of the Species.
''Bulletin of the National Museum of Nature and Science, Ser. A., 40 (3): 141-160.''
* '' Drombus'' * '' Ebomegobius'' * '' Echinogobius'' * '' Economidichthys'' * '' Egglestonichthys'' * '' Ego'' * ''
Elacatinus ''Elacatinus'' is a genus of small marine gobies, often known collectively as the neon gobies. Although only one species, ''E. oceanops'', is technically the "neon goby," because of their similar appearance, other members of the genus are genera ...
'' * '' Eleotrica'' * '' Evermannia'' * '' Eviota'' * '' Exyrias'' * '' Favonigobius'' * '' Feia'' * '' Fusigobius'' * '' Gammogobius'' * '' Ginsburgellus'' * '' Gladiogobius'' * ''
Glossogobius ''Glossogobius'' is a genus of gobies native to fresh, brackish and marine waters from Africa to the coasts of the western Pacific Ocean. They are found in Madagascar, South Africa, Japan, Thailand, Australia, Indonesia, Bangladesh, the Philippin ...
'' * ''
Gobiodon ''Gobiodon'' is a genus of gobies also known as coral gobies or "clown gobies" (which can also mean the related genus '' Microgobius''). Generally, coral gobies, unlike the rest of the family Gobiidae, are not burrowers, but instead prefer to in ...
'' * '' Gobiopsis'' * '' Gobiosoma'' Tornabene, L. & Van Tassell, J.L. (2014): Redescription of the goby genus ''Gobiosoma'' (Teleostei: Gobiidae: Gobiosomatini), with the synonymy of the genus ''Enypnias''. ''Journal of Natural History, 48 (23-24): 1413-1437.'' * ''
Gobius ''Gobius'' is a genus of fish in the family Gobiidae native to fresh, brackish and marine waters of and around Europe, Africa and Asia. It contains the typical gobies, being the type genus of the formerly recognised subfamily Gobiinae and famil ...
'' * '' Gobiusculus'' * '' Gobulus'' * '' Gorogobius'' * '' Grallenia'' * '' Gymneleotris'' * '' Hazeus'' * ''
Hetereleotris ''Hetereleotris'' is a genus of gobies native to the western Indian Ocean and the western Pacific Ocean. Species There are currently 18 recognized species in this genus: * '' Hetereleotris apora'' Hoese & R. Winterbottom, 1979 (Poreless goby) ...
'' * '' Heterogobius'' * '' Heteroplopomus'' * '' Hyrcanogobius'' * '' Istigobius'' * '' Kelloggella'' * '' Knipowitschia'' * '' Koumansetta'' * ''
Larsonella ''Larsonella pumila'' is a species of goby native to the Indian Ocean from the coast of Africa to the western Pacific Ocean. This species grows to a length of SL. This species is the only known member of its genus. The specific name honours ...
'' * '' Lebetus'' * '' Lesueurigobius'' * '' Lobulogobius'' * '' Lophiogobius'' * ''
Lophogobius ''Lophogobius'' is a small genus of gobies native to the Pacific and western Atlantic oceans. Species There are currently 2 recognized species in this genus: * '' Lophogobius cristulatus'' Ginsburg, 1939 * '' Lophogobius cyprinoides'' (Pallas ...
'' * '' Lotilia'' * '' Lubricogobius'' * '' Luposicya'' * ''
Lythrypnus ''Lythrypnus'' is a genus of gobies native to the Atlantic and Pacific coasts of the Americas including Cocos Island and the Galapagos Islands. Species There are currently 20 recognized species in this genus: * '' Lythrypnus alphigena'' W. A. ...
'' * '' Macrodontogobius'' * '' Mahidolia'' * '' Mangarinus'' * '' Mauligobius'' * ''
Mesogobius ''Mesogobius'' is one of the genera of benthophiline gobiid fishes native to the basins of the Black and Caspian Seas. Species There are two or three recognized species in this genus:.Brian W. CoadFreshwater Fishes of Iran(accessed 18 Feb 201 ...
'' * '' Microgobius'' * '' Minysicya'' * '' Myersina'' * '' Nematogobius'' * ''
Neogobius ''Neogobius'' is a genus of goby native to Black Sea and the Caspian Sea basins. It is part of the broader Benthophilinae subfamily which is also endemic to the same region. Nevertheless, two ''Neogobius'' species have recently turned out to be h ...
'' * ''
Nes The Nintendo Entertainment System (NES) is an 8-bit third-generation home video game console produced by Nintendo. It was first released in Japan in 1983 as the commonly known as the The NES, a redesigned version, was released in American ...
'' * '' Nesogobius'' * '' Obliquogobius'' * '' Odondebuenia'' * '' Ophiogobius'' * '' Oplopomops'' * '' Oplopomus'' * ''
Opua Opua is a locality in the Bay of Islands, in the sub-tropical Northland Region of New Zealand. It is notable as the first port for overseas yachts arriving in the country after crossing the Pacific Ocean. In the original 1870s plans for the ...
'' * ''
Padogobius ''Padogobius'' is a genus of fish in the family Gobiidae, the gobies. They are native to fresh waters of southern Europe. Species There are currently two recognized species in this genus: * ''Padogobius bonelli'' ( Bonaparte, 1846) (Padanian gob ...
'' * '' Palatogobius'' * '' Palutrus'' * '' Parachaeturichthys'' * '' Paragobiodon'' * '' Paratrimma'' * '' Pariah'' * '' Parkraemeria'' * '' Parrella'' * '' Pascua'' * '' Phoxacromion'' * '' Phyllogobius'' * '' Platygobiopsis'' * '' Pleurosicya'' * '' Polyspondylogobius'' * ''
Pomatoschistus ''Pomatoschistus'' is a genus of gobies native to fresh, brackish and marine waters of Europe, the eastern Atlantic Ocean and the Mediterranean Sea. Species There are currently 13 recognized species in this genus: * '' Pomatoschistus anatoliae' ...
'' * ''
Ponticola ''Ponticola'' is a genus of gobies native mostly to fresh waters of the Black Sea - Caspian Sea region in Eurasia. Some species occur in the brackish-water Black and Caspian seas themselves. It was considered to be part of the broader goby subfam ...
'' * '' Porogobius'' * '' Priolepis'' * ''
Proterorhinus ''Proterorhinus'' is a genus of fishes, known as the tubenose gobies. These gobies, gobiid fish are native to Eurasia where they occur in the region of the Caspian Sea, Caspian and Black Sea, Black seas, inhabiting marine, brackish and fresh wate ...
'' * '' Psammogobius'' * '' Pseudaphya'' * '' Psilogobius'' * '' Psilotris'' * '' Pycnomma'' * '' Rhinogobiops'' * '' Risor'' * '' Robinsichthys'' * '' Signigobius'' * '' Silhouettea'' * '' Siphonogobius'' * '' Speleogobius'' * '' Stonogobiops'' * '' Sueviota'' * '' Sufflogobius'' * '' Thorogobius'' * '' Tigrigobius'' * '' Tomiyamichthys'' * '' Trimma'' * '' Trimmatom'' * '' Tryssogobius'' * ''
Valenciennea ''Valenciennea'' is a genus of bottom-dwelling, small, bottom-dwelling fish in the family (biology), family Gobiidae. They are found over Benthic zone, sandy bottoms, often at coral reefs in the Indo-Pacific. The members of the genus tend to rest ...
'' * ''Vanderhorstia'' * ''Vanneaugobius'' * ''Varicus'' * ''Vomerogobius'' * ''Wheelerigobius'' * ''Yoga (fish), Yoga'' * ''Yongeichthys'' * ''Zebrus'' * ''Zosterisessor''


References

Gobiinae, Gobiidae Taxa named by Georges Cuvier {{Gobiidae-stub