Trox Scaber
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''Trox scaber'' is a
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
of the family
Trogidae Trogidae, sometimes called hide beetles, is a family of beetles with a distinctive warty or bumpy appearance. Found worldwide, the family includes about 300 species contained in four or five genera. Trogids range in length from 2 to 20 mm. Thei ...
. The 5 to 8 mm long insect is found worldwide, including in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
, and lives in bird nests.Norman H. Joy, 1932 ''
A Practical Handbook of British Beetles ''A Practical Handbook of British Beetles'' is a two-volume work on the British beetle fauna, by Norman H. Joy, first published by H. F. & G. Witherby in January 1932. Contents Volume one (xxviii + 622 pages) consists of the text (largely a ...
''
The elytra and pronotum are roughly sculptured, and there are bristly scales on the elytral ridges. The larvae develop in tawny owl and other owl nests but also in the hollow tree nests of jackdaw,
woodpecker Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extreme polar regions. ...
and others. They feed mainly on small dry carcasses, especially when dry.


Name and classification

''Trox scaber'' was previously put in the Scarabaeidae because of the similarity of its antennae to that of members of this family. The genus ''Trox'' has 46 species throughout the world. In Europe, there are 21 species, all belonging to the subgenus ''Trox''. There are about 8 in Central Europe. The name ''Trox'' (from the Ancient Greek τροξ, meaning "chewer") may have been given because the mandible is forked. It is described by Fabricius as ''Maxilla bifida''. The species name ''scaber'' (Latin ''scaber'' = rough) alludes to the rough surface of the elytra.


Appearance

The beetle is brown to black-brown. The antennae are rust red. The bristles are brownish-yellow and not black. The body is short, obovate and dome-shaped. The head is recessed into the pronotum. The antennae have 10 segments. The last 3 segments form a fan and, because of the fine hairs, are not shiny. The first flagellomere is long and slightly undermined underneath. The
mouthparts Mouthparts may refer to: * The parts of a mouth ** Arthropod mouthparts *** Insect mouthparts {{disambig ...
point downwards and are not level with the clypeus. The mandible is short, with small, nearly smooth masticatory processes. The maxillary palpi are rather short, and the terminal segment is elongated. The third segment of the labial palpus is rounded. The lateral and basal margins of the pronotum are bordered. The sides are edged with short bristles. At the base are less regular, somewhat longer bristles. The curved disc is coarsely sculpted and also has rows of bristles. The elytra cover the
telson The telson () is the posterior-most division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment of the arthropod body, or an additional division that is not a true segment on accou ...
. Each elytron bears 10 thin, even and shallowly punctured stripes. Between these are alternatingly about four times as wide, higher arched and about three times as wide, less curved spaces. On each of the higher, wider surfaces is a row of small humps, which are occupied with short, strong bristles. On the narrower sections, there are no humps and the bristles are much sparser and found as a row of very small tufts of bristles. The sides of the elytra are bordered with a row of short bristles. Between the forelegs, the pronotum is posteriorly extended. This prosternal process is short and pointy. Only 5 sternites can be found on the abdomen. The coxae of the middle legs are very small and round. The
legs A leg is a weight-bearing and locomotive anatomical structure, usually having a columnar shape. During locomotion, legs function as "extensible struts". The combination of movements at all joints can be modeled as a single, linear element ...
have five-segmented tarsi. The outer edges of the tibiae of the front legs are serrated, which hints that they are fossorial legs.


Living habits

The heat-loving beetles can make chirping noises by rubbing the abdomen on the elytra. The species is mainly found in the substructure of older large birds' nests, but are also found in smaller bird nests, bird carcasses and hides, which they gnaw on. Occasionally, they are also found at the entrance of underground animal dens. The adults fly only grudgingly.


Distribution

The species is found almost worldwide. In Central Europe, they are common in low-elevation terrain and are rare to the
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Sw ...
. File:Trox scaber bl.jpg, head File:Trox scaber bl2.jpg, dorsal File:Trox scaber side.jpg, side File:Trox scaber underside.jpg, underside


Literature

* Heinz Freude, Karl Wilhelm Harde, Gustav Adolf Lohse (Hrsg.): ''Die Käfer Mitteleuropas''. Band 8. ''Teredilia Heteromera Lamellicornia''. Elsevier, Spektrum, Akademischer Verlag, München 1969, . * Gustav Jäger (Hrsg.): '' C. G. Calwer's Käferbuch''. K. Thienemanns, Stuttgart 1876, 3. Auflage. * Klaus Koch: ''Die Käfer Mitteleuropas Ökologie''. 1. Auflage. Band 2. Goecke & Evers, Krefeld 1989, .


References

J.C. Fabricius: ''Systema entomologiae, sistens insectorum classes...'' Flensburgi, Lipsiae 177
Original description of genus Trox at GDZ
Fauna Europaea
''Trox'' in BioLib
Sigmund Schenkling
''Erklärung der wissenschaftlichen Käfernamen (Art)''
(Eng. "Explanation of scientific names of beetles (species)")
Sigmund Schenkling
''Erklärung der wissenschaftlichen Käfernamen (Gattung)''
(Eng. "Explanation of scientific names of beetles (genus)")
{{Taxonbar, from=Q143275 scaber Beetles described in 1767