Tropidophorus Beccarii
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''Tropidophorus beccarii'', also known commonly as Beccari's keeled skink and Beccari's water skink, is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
in the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Lygosominae Lygosominae is the largest subfamily of skinks in the family Scincidae. The subfamily can be divided into a number of genus groups. If the rarely used taxonomic rank of infrafamily is employed, the genus groups would be designated as such, but s ...
of the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Scincidae. The species is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to the island of
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and eas ...
.


Etymology

The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, ''beccarii'', is in honor of Italian botanist
Odoardo Beccari Odoardo Beccari (16 November 1843 – 25 October 1920) was an Italian botanist famous for his discoveries in Indonesia, particularly New Guinea, and Australia. He has been called the greatest botanist to ever study Malesia. His author abbreviat ...
.


Geographic range

On the island of Borneo, ''T. beccarii'' is found in
Brunei Brunei ( , ), formally Brunei Darussalam ( ms, Negara Brunei Darussalam, Jawi alphabet, Jawi: , ), is a country located on the north coast of the island of Borneo in Southeast Asia. Apart from its South China Sea coast, it is completely sur ...
, northern
Kalimantan Kalimantan () is the Indonesian portion of the island of Borneo. It constitutes 73% of the island's area. The non-Indonesian parts of Borneo are Brunei and East Malaysia. In Indonesia, "Kalimantan" refers to the whole island of Borneo. In 2019, ...
(
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
),
Sabah Sabah () is a States and federal territories of Malaysia, state of Malaysia located in northern Borneo, in the region of East Malaysia. Sabah borders the Malaysian state of Sarawak to the southwest and the North Kalimantan province of Indone ...
, and
Sarawak Sarawak (; ) is a States and federal territories of Malaysia, state of Malaysia. The largest among the 13 states, with an area almost equal to that of Peninsular Malaysia, Sarawak is located in northwest Borneo Island, and is bordered by the M ...
(
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
).


Habitat

The preferred natural
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s of ''T. beccarii'' are
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
and freshwater
wetlands A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The ...
, at altitudes from sea level to .


Description

Dorsally Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
, ''T. beccarii'' is reddish brown, with dark brown crossbands. The flanks have whitish spots. Ventrally, it is yellowish. ''T. beccarii'' may attain a snout to vent length (SVL) of . The largest specimen measured by
Boulenger Boulenger is a surname. Notable people with the surname include: * Benjamin Boulenger (born 1990), French footballer * Edward George Boulenger (1888–1946), British zoologist, director of aquarium at London Zoo * George Albert Boulenger (1858–1 ...
had an SVL of , and a tail length of . Boulenger GA (1887). ''Catalogue of the Lizards in the British Museum (Natural History). Second Edition. Volume III. ... Scincidæ ....'' London: Trustees of the British Museum (Natural History). (Taylor and Francis, printers). xii + 575 pp. + Plates I–XL. (''Tropidophorus beccarii'', pp. 360–361).


Diet

''T. beccarii''
preys Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
upon water
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s. Das I (2006). ''A Photographic Guide to Snakes and other Reptiles of Borneo''. Sanibel Island, Florida: Ralph Curtis Books. 144 pp. . (''Tropidophorus beccarii'', p. 118).


Reproduction

''T. beccarii'' is
viviparous Among animals, viviparity is development of the embryo inside the body of the parent. This is opposed to oviparity which is a reproductive mode in which females lay developing eggs that complete their development and hatch externally from the m ...
. Average
litter Litter consists of waste products that have been discarded incorrectly, without consent, at an unsuitable location. Litter can also be used as a verb; to litter means to drop and leave objects, often man-made, such as aluminum cans, paper cups, ...
size is four.


References


Further reading

* Goldberg SR (2017). "Reproduction in Beccari's Keeled Skinks, ''Tropidophorus beccarii'' (Squamata: Scincidae), from Borneo". ''Bulletin of the Chicago Herpetological Society'' 52: 65. * Hikida T, Ota H (1994). "''Sphenomorphus aquaticus'' Malkmus, 1991, a junior synonym of ''Tropidophorus beccarii'' (Peters, 1871) (Reptilia: Squamata: Scincidae)". ''Bonner Zoologische Beiträge'' 45 (1): 57–60. * Peters W (1871). "''Über neue Reptilien aus Ostafrika und Sarawak (Borneo), vorzüglich aus der Sammlung des Hrn. Marquis J. Doria zu Genua''". ''Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin'' 1871: 566–581. (''Tropidophorus beccarii'', new species, p. 574). (in German). beccarii Endemic fauna of Borneo Reptiles of Brunei Reptiles of Indonesia Reptiles of Malaysia Reptiles described in 1871 Taxa named by Wilhelm Peters Reptiles of Borneo {{Lygosominae-stub