Tropidoclonion
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''Tropidoclonion'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
snake Snakes are elongated, Limbless vertebrate, limbless, carnivore, carnivorous reptiles of the suborder Serpentes . Like all other Squamata, squamates, snakes are ectothermic, amniote vertebrates covered in overlapping Scale (zoology), scales. Ma ...
in the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Natricinae of the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Colubridae Colubridae (, commonly known as colubrids , from la, coluber, 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest species of the family date back to the Oligocene epoch. Colubrid snakes are found on ever ...
. The genus is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
, containing the sole
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
''Tropidoclonion lineatum'', commonly known as the lined snake. The species is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
.


Common names

Additional common names for ''T. lineatum'' include common snake, dwarf garter snake, grass snake, line snake, ribbon snake, streaked snake, striped snake, and swamp snake.


Subspecies

Four
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognized as being valid, including the
nominotypical subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
. *''Tropidoclonion lineatum annectens'' – central lined snake *''Tropidoclonion lineatum lineatum'' – northern lined snake *''Tropidoclonion lineatum mertensi'' – Mertens' lined snake *''Tropidoclonion lineatum texanum'' – Texas lined snake ''
Nota bene (, or ; plural form ) is a Latin phrase meaning "note well". It is often abbreviated as NB, n.b., or with the ligature and first appeared in English writing . In Modern English, it is used, particularly in legal papers, to draw the atten ...
'': A trinomial authority in parentheses indicates that the subspecies was originally described in a genus other than ''Tropidoclonion''.


Etymology

The subspecific name, ''mertensi'', is in honor of German herpetologist
Robert Mertens Robert Friedrich Wilhelm Mertens (1 December 1894 – 23 August 1975) was a German herpetologist. Several taxa of reptiles are named after him.Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore ...
.


Geographic range

The lined snake is found throughout the central
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
from
Illinois Illinois ( ) is a U.S. state, state in the Midwestern United States, Midwestern United States. Its largest metropolitan areas include the Chicago metropolitan area, and the Metro East section, of Greater St. Louis. Other smaller metropolita ...
to
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
.


Habitat

The preferred
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
of ''T. lineatum'' is
grassland A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...
areas with soft, moist soils.


Description

The lined snake is olive
green Green is the color between cyan and yellow on the visible spectrum. It is evoked by light which has a dominant wavelength of roughly 495570 Nanometre, nm. In subtractive color systems, used in painting and color printing, it is created by ...
to
brown Brown is a color. It can be considered a composite color, but it is mainly a darker shade of orange. In the CMYK color model used in printing or painting, brown is usually made by combining the colors orange and black. In the RGB color model used ...
with a distinctive tan or
yellow Yellow is the color between green and orange on the spectrum of light. It is evoked by light with a dominant wavelength of roughly 575585 nm. It is a primary color in subtractive color systems, used in painting or color printing. In the R ...
stripe running down the middle of the back from head to
tail The tail is the section at the rear end of certain kinds of animals’ bodies; in general, the term refers to a distinct, flexible appendage to the torso. It is the part of the body that corresponds roughly to the sacrum and coccyx in mammals, r ...
. It has similar stripes, one down each side on scale rows 2 and 3. On the belly, it has a double row of clean-cut black half-moon spots running down the middle. Conant R (1975). ''A Field Guide to Reptiles and Amphibians of Eastern and Central North America, Second Edition''. Boston: Houghton Mifflin. xviii + 429 pp. + Plates 1-48. (hardcover), (paperback). (''Tropidoclonion lineatum'', pp. 166-167 + Plate 24 + Map 123). It has a narrow head and small
eye Eyes are organs of the visual system. They provide living organisms with vision, the ability to receive and process visual detail, as well as enabling several photo response functions that are independent of vision. Eyes detect light and conv ...
s. Adult size is typically less than 35 cm (14 inches) in total length (including tail). However, maximum recorded total length is 53 cm (21 in). Smith HM, Brodie ED Jr (1982). ''Reptiles of North America: A Guide to Field Identification''. New York: Golden Press. 240 pp. (paperback). (''Tropidoclonion lineatum'', pp. 152-153). The keeled
dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...
are arranged in 19 rows at midbody. There are only 5 or 6
upper labials In reptiles, the supralabial scales, also called upper-labials, are those scales that border the mouth opening along the upper jaw. They do not include the median scaleWright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates ( ...
.


Behavior

The lined snake is semi
fossorial A fossorial () animal is one adapted to digging which lives primarily, but not solely, underground. Some examples are badgers, naked mole-rats, clams, meerkats, and mole salamanders, as well as many beetles, wasps, and bees. Prehistoric eviden ...
, spending most of its time hiding under rocks, leaf litter, logs, or buried in the soil.


Diet

The majority of the diet of ''T. lineatum'' consists of
earthworm An earthworm is a terrestrial invertebrate that belongs to the phylum Annelida. They exhibit a tube-within-a-tube body plan; they are externally segmented with corresponding internal segmentation; and they usually have setae on all segments. Th ...
s.


Reproduction

The lined snake is
ovoviviparous Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a term used as a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop insi ...
, the young being born in August. The average brood is seven or eight.Force ER (1931). "Habits and Birth of the Lined Snake, ''Tropidoclonion lineatum'' (Hallowell)". ''Copeia'' 1931: 51-53. The newborn juveniles are 10–12 cm (4-4¾ in.) long at birth.


References


Further reading

* Behler JL, King FW (1979). ''The Audubon Society Field Guide to North American Reptiles and Amphibians''. New York: Alfred A. Knopf. 743 pp.657 color plates. . (''Tropidoclonion lineatum'', pp. 677-678 + Plate 507). * Conant R, Bridges W (1939). ''What Snake Is That? A Field Guide to the Snakes of the United States East of the Rocky Mountains''. (With 108 drawings by Edmond Malnate). New York and London: D. Appleton-Century. Frontispiece map + viii + 163 pp. + Plates A-C, 1-32. (''Tropidoclonion lineatum'', pp. 114–115 + Plate 21, Figure 63). * Hallowell E (1856). "Notice of a Collection of Reptiles from Kansas and Nebraska, presented to the Academy of Natural Sciences, by Dr. Hammond, U. S. A." ''Proc. Acad. Nat. Sci. Philadelphia'' 8: 238-253. (''Microps'', new genus, p. 240; ''Microps lineatus'', new species, p. 241). * Powell R, Conant R, Collins JT (2016). ''Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America, Fourth Edition''. Boston and New York: Houghton Mifflin Harcourt. xiv + 494 pp., 47 plates, 207 figures. . (''Tropidoclonion lineatum'', p. 433 + Plate 44). * Stebbins RC (2003). ''A Field Guide to Western Reptiles and Amphibians, Third Edition''. The Peterson Field Guide Series ®. Boston and New York: Houghton Mifflin. xiii + 533 pp. (paperback). (''Tropidoclonion lineatum'', p. 391 + Plate 50 + Map 168). * Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Ithaca and London: Comstock Publishing Associates, a Division of Cornell University Press. 1,105 pp. (in 2 volumes) (Genus ''Tropidoclonion'' p. 879 + Figure 20, a-c, on p. 71; and species ''Tropidoclonion lineatum'', pp. 879–884, Figure 252, Map 62). * Zim HS, Smith HM (1956). ''Reptiles and Amphibians: A Guide to Familiar American Species: A Golden Nature Guide''. Revised Edition. New York: Simon and Schuster. 160 pp. ("Lined Snake", pp. 106, 156).


External links

*
''Tropidoclonion'' at Oklahoma SnakesLined Snake
Reptiles and Amphibians of Iowa {{Taxonbar, from=Q5082984 Colubrids Reptiles of the United States Extant Pleistocene first appearances Monotypic snake genera Taxa named by Edward Drinker Cope