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Tritheledontidae, the tritheledontids or ictidosaurs, is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of small to medium-sized (about 10 to 20 cm long)
cynodont The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety ...
s. They were highly
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
-like, specialized cynodonts, although they still retained a few
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
-like anatomical traits. Tritheledontids were mainly
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
or
insectivorous A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant that eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores were ...
, though some species may have developed
omnivory An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nut ...
. Their skeletons show that they had a close relationship to mammals. Tritheledontids or their closest relatives may have given rise to the
mammaliaforms Mammaliaformes ("mammalian forms") is a clade that contains the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts. It is defined as the clade originating from the most recen ...
. The tritheledontids were one of the longest lived non-mammalian
therapsid Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more ...
lineages, living from the late
Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...
to the
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
period. Tritheledontids became extinct in the Jurassic period, possibly due to competition with
prehistoric mammal This is an incomplete list of prehistoric mammals. It does not include extant mammals or recently extinct mammals. For extinct primate species, see: list of fossil primates.Mikko's Phylogeny Archiv Mammaliaformes ' *Genus †'' Adelobasileus ...
s such as the
eutriconodont Eutriconodonta is an order of early mammals. Eutriconodonts existed in Asia, Africa, Europe, North and South America during the Jurassic and the Cretaceous periods. The order was named by Kermack ''et al.'' in 1973 as a replacement name for the ...
s. They are known from finds in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
and
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
, indicating that they may have lived only on the
supercontinent In geology, a supercontinent is the assembly of most or all of Earth's continent, continental blocks or cratons to form a single large landmass. However, some geologists use a different definition, "a grouping of formerly dispersed continents", ...
of
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
. The family Tritheledontidae was named by South African paleontologist
Robert Broom Robert Broom FRS FRSE (30 November 1866 6 April 1951) was a British- South African doctor and palaeontologist. He qualified as a medical practitioner in 1895 and received his DSc in 1905 from the University of Glasgow. From 1903 to 1910, he ...
in 1912. May be downloaded from: https://archive.org/details/annalsofsouthafr07sout The family is often misspelled "Trithelodontidae". It is possible that tritheledontids had
vibrissae Vibrissae (; singular: vibrissa; ), more generally called Whiskers, are a type of stiff, functional hair used by mammals to sense their environment. These hairs are finely specialised for this purpose, whereas other types of hair are coarser ...
, according to the
PBS The Public Broadcasting Service (PBS) is an American public broadcasting, public broadcaster and Non-commercial activity, non-commercial, Terrestrial television, free-to-air television network based in Arlington, Virginia. PBS is a publicly fu ...
documentary, ''Your Inner Fish''. A common ancestor of all therian mammals did so. It is possible that the development of the whisker sensory system played an important role in mammalian development, more generally.


Genera

* '' Chaliminia'' * ''
Diarthrognathus ''Diarthrognathus'' ("Two joint jaw") is an extinct genus of tritheledontid cynodonts, known from fossil evidence found in South Africa and first described in 1958 by A.W. Crompton.Rieppel, Olivier. Evolutionary Theory and the Creation Controvers ...
'' * '' Elliotherium'' * '' Irajatherium'' * ''
Pachygenelus ''Pachygenelus'' is an extinct genus of tritheledontid cynodonts. Fossils have been found from the Karoo basin in South Africa and date back to the Early Jurassic. ''Pachygenelus'' had both an articular- quadrate and dentary-squamosal jaw joint ...
'' * ''
Riograndia ''Riograndia'' is an extinct genus of tritheledontid cynodonts from the Late Triassic of South America. The type and only species is ''Riograndia guaibensis''. Remains have been found in the Caturrita Formation of the geopark of Paleorrota. I ...
'' * '' Tritheledon''


See also

*
Evolution of mammals The evolution of mammals has passed through many stages since the first appearance of their synapsid ancestors in the Pennsylvanian sub-period of the late Carboniferous period. By the mid-Triassic, there were many synapsid species that looked l ...


References


External links


PBS YouTube animated video
of a tritheledontid {{Taxonbar, from1=Q21646750, from2=Q140837 Late Triassic first appearances Middle Jurassic extinctions Prehistoric therapsid families