Trioceros Johnstoni
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''Trioceros johnstoni'', known commonly as Johnston's chameleon, Johnston's three-horned chameleon, and the Ruwenzori three-horned chameleon, is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
chameleon Chameleons or chamaeleons (family Chamaeleonidae) are a distinctive and highly specialized clade of Old World lizards with 202 species described as of June 2015. The members of this family are best known for their distinct range of colors, bein ...
, a
lizard Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Chamaeleonidae. The species is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to highlands in the
Albertine Rift The Albertine Rift is the western branch of the East African Rift, covering parts of Uganda, the Democratic Republic of the Congo (DRC), Rwanda, Burundi and Tanzania. It extends from the northern end of Lake Albert to the southern end of Lake Tan ...
in central Africa. It reaches up to in total length (including tail). Only the adult male has three horns. The female is hornless.


Taxonomy and etymology

The three long annulated horns in the male ''T. johnstoni'' makes it superficially similar to '' T. jacksonii'' and '' T. werneri'', and to an extent also to the short-horned '' T. fuelleborni'' and the smooth-horned '' T. oweni'', but they are not close relatives. Its nearest relative is the hornless '' T. ituriensis''. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''johnstoni'' was given in honour of the British explorer
Harry Johnston Sir Henry Hamilton Johnston (12 June 1858 – 31 July 1927), known as Harry Johnston, was a British explorer, botanist, artist, colonial administrator, and linguist who travelled widely in Africa and spoke many African languages. He publishe ...
. Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore: Johns Hopkins University Press. xiii + 296 pp. . (''Trioceros johnstoni'', p. 135).


Distribution and habitat

''T. johnstoni'' is found in forests at altitudes between in the Albertine Rift of eastern
Democratic Republic of the Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...
, western
Burundi Burundi (, ), officially the Republic of Burundi ( rn, Repuburika y’Uburundi ; Swahili language, Swahili: ''Jamuhuri ya Burundi''; French language, French: ''République du Burundi'' ), is a landlocked country in the Great Rift Valley at the ...
, western
Rwanda Rwanda (; rw, u Rwanda ), officially the Republic of Rwanda, is a landlocked country in the Great Rift Valley of Central Africa, where the African Great Lakes region and Southeast Africa converge. Located a few degrees south of the Equator ...
, and southwestern
Uganda }), is a landlocked country in East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territor ...
, but also tolerates semi-urbanized environments as long as some trees and bushes remain.


Behaviour

Males of ''T. johnstoni'' are fiercely
territorial A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
and readily will fight other males, using their horns and biting.


Reproduction

''T. johnstoni'' is
oviparous Oviparous animals are animals that lay their eggs, with little or no other embryonic development within the mother. This is the reproductive method of most fish, amphibians, most reptiles, and all pterosaurs, dinosaurs (including birds), and ...
, with the female laying 4–23 eggs per clutch.


References


Further reading

* Spawls S, Howell K, Hinkel H, Menegon M (2018). ''Field Guide to East African Reptiles, Second Edition''. London: Bloomsbury Wildlife. 624 pp. . (''Trioceros johnstoni'', pp. 293–294). * Tilbury CR, Tolley KA (2009). "A re-appraisal of the systematics of the African genus ''Chamaeleo'' (Reptilia: Chamaeleonidae)". ''Zootaxa'' 2079: 57–68. (''Trioceros johnstoni'', new combination). Trioceros Lizards of Africa Reptiles described in 1901 Taxa named by George Albert Boulenger {{chameleon-stub