''Protobothrops kaulbacki'',
commonly known as the Kaulback's lance-headed pitviper
[ or Kaulback's lance-headed pit viper,][Gumprecht A, Tillack F, Orlov NL, ]Captain A
Captain is a title, an appellative for the commanding officer of a military unit; the supreme leader of a navy ship, merchant ship, aeroplane, spacecraft, or other vessel; or the commander of a port, fire or police department, election precinct, e ...
, Ryabov S (2004). ''Asian Pit vipers''. Berlin: Geitje Books. First Edition. 368 pp. . is a venomous
Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
pit viper species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
endemic
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to Asia
Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
. No subspecies are currently recognized.
Etymology
The specific name, ''kaulbacki'', is in honor of British explorer Ronald Kaulback
Ronald John Henry Kaulback (23 July 1909 – 2 October 1995) was a British explorer, botanist and geographer. He was the son of Colonel H. A. Kaulback and had a younger brother Roy (known as Bill). After studying at the University of Cambridge and ...
.[
]
Description
Adult males of ''P. kaulbacki'' may attain a total length of , which includes a tail long. Females may grow longer: maximum total length , tail . Dorsally
Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
, it is green, with a vertebral series of dark angular spots, which may be joined to form a zigzag stripe. The top of the head is black with yellow stripes. Ventrally, except for the whitish throat area, it is gray with large squarish or crescent-shaped yellow spots.[ Smith MA (1943). ''The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. III.—Serpentes''. London: Secretary of State for India. (Taylor and Francis, printers). xii + 583 pp. (''Protobothrops kaulbacki'', p. 512).] Scalation includes 25 rows of dorsal scales
In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales
In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down t ...
at midbody, 201–212 ventral scales
In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...
, 66–78 subcaudal scales, and 8 supralabial scales of which the third is the largest.[
]
Geographic range
''Protobothrops kaulbacki'' had originally been known only from the type locality, which is "Pangnamdim, north of the Triangle, Upper Burma
Upper Myanmar ( my, အထက်မြန်မာပြည်, also called Upper Burma) is a geographic region of Myanmar, traditionally encompassing Mandalay and its periphery (modern Mandalay, Sagaing, Magway Regions), or more broadly speak ...
" (Myanmar
Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
).[ In 2005 it was reported also from ]Tibet
Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa, Taman ...
(China).[ Rao D, Zhao E (2005). "A New Record from China – ''Protobothrops kaulbacki'' (Reptilia, Serpentes, Viperidae)". ''Acta Zootaxonomica Sinica'' 30: 209-211.] It also occurs in Arunachal Pradesh
Arunachal Pradesh (, ) is a state in Northeastern India. It was formed from the erstwhile North-East Frontier Agency (NEFA) region, and became a state on 20 February 1987. It borders the states of Assam and Nagaland to the south. It shares ...
(NE India).[
]
Reproduction
''Protobothrops kaulbacki'' is oviparous. The adult female lays a clutch of 6–32 egg
An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...
s in a hole in the ground, and then remains with the eggs to guard them. Eggs measure 48–53 mm x 26–27 mm (about 2 in x 1 in). Each hatchling is 26–27 cm (about 10½ inches) in total length.
References
Further reading
*Kraus, Fred; Mink, Daniel G.; Brown, Wesley M. (1996). "Crotaline Intergeneric Relationships Based on Mitochondrial DNA Sequence Data". ''Copeia'' 1996 (4): 763–773. (''Protobothrops kaulbacki'', new combination).
* Smith MA (1940). "The Amphibians and Reptiles obtained by Mr. Ronald Kaulback in Upper Burma". ''Records of the Indian Museum'' 42: 465–486. (''Protobothrops kaulbacki'', new species, p. 485, Plate VIII, Figure 5).
kaulbacki
Snakes of China
Reptiles of India
Reptiles of Myanmar
Fauna of Tibet
Reptiles described in 1940
Taxa named by Malcolm Arthur Smith
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