Triglops Macellus
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''Triglops'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...
es belonging to the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Cottidae The Cottidae are a family of fish in the superfamily Cottoidea, the sculpins. It is the largest sculpin family, with about 275 species in 70 genera.Kane, E. A. and T. E. Higham. (2012)Life in the flow lane: differences in pectoral fin morphology ...
, the typical
sculpin A sculpin is a type of fish that belongs to the superfamily Cottoidea in the order Scorpaeniformes.Kane, E. A. and T. E. Higham. (2012)Life in the flow lane: differences in pectoral fin morphology suggest transitions in station-holding demand a ...
s. These fishes are found in the North Pacific, Arctic and North Atlantic Oceans.


Taxonomy

''Triglops'' was first proposed as a monospecific subgenus of '' Cottus'' in 1830 by the Danish
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...
Johan Reinhardt when he described ''Triglops pingelii'', giving its type locality as Quanneoen, south of Frederikshaab in
western Greenland Kitaa, originally Vestgrønland ("West Greenland"), is a former administrative division of Greenland. It was by far the most populated of the divisions, being home to almost 90% of the total population. The divisions were de facto replaced by st ...
. The 5th edition of ''
Fishes of the World ''Fishes of the World'' by the American ichthyologist Joseph S. Nelson (1937–2011) is a standard reference for fish systematics. Now in its fifth edition (2016), the work is a comprehensive overview of the diversity and classification of the ...
'' classifies the genus ''Triglops'' within the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Cottinae Cottinae is a subfamily of ray-finned fishes belonging to the family Cottidae, the typical sculpin A sculpin is a type of fish that belongs to the superfamily Cottoidea in the order Scorpaeniformes.Kane, E. A. and T. E. Higham. (2012)Life in t ...
of the family Cottidae, however, other authors classify the genus within the subfamily Triglopinae of the family
Psychrolutidae The fish family Psychrolutidae (commonly known as blobfishes, toadfishes, flathead sculpins, tadpole sculpins,) contains over 35 recognized species in 8 genera. This family consists of bottom-dwelling marine sculpins shaped like tadpoles, with l ...
.


Species

''Triglops'' currently contains 10 recognized species in this genus: * '' Triglops dorothy'' Pietsch & J. W. Orr, 2006 (Dorothy's sculpin) * '' Triglops forficatus'' (
Gilbert Gilbert may refer to: People and fictional characters * Gilbert (given name), including a list of people and fictional characters *Gilbert (surname), including a list of people Places Australia * Gilbert River (Queensland) * Gilbert River (South ...
, 1896)
(Scissortail sculpin) * '' Triglops jordani'' (
Schmidt Schmidt may refer to: * Schmidt (surname), including list of people with the surname * Schmidt (singer) (born 1990), German pop and jazz singer * Schmidt (lunar crater), a small lunar impact crater * Schmidt (Martian crater), a List of craters on ...
, 1904)
(Jordan's sculpin) * '' Triglops macellus'' ( T. H. Bean, 1884) (Roughspine sculpin) * ''
Triglops metopias ''Triglops'' is a genus of marine ray-finned fishes belonging to the Family (biology), family Cottidae, the typical sculpins. These fishes are found in the North Pacific, Arctic and North Atlantic Oceans. Taxonomy ''Triglops'' was first proposed ...
'' Gilbert &
Burke Burke is an Anglo-Norman Irish surname, deriving from the ancient Anglo-Norman and Hiberno-Norman noble dynasty, the House of Burgh. In Ireland, the descendants of William de Burgh (–1206) had the surname ''de Burgh'' which was gaelicised ...
, 1912
(Alaskan sculpin) * ''
Triglops murrayi ''Triglops murrayi'', the moustache sculpin, is a species of marine ray-finned fish belonging to the Family (biology), family Cottidae, the typical sculpins. This fish is found in the North Atlantic Ocean. Taxonomy ''Triglops murrayi'' was first ...
'' Günther, 1888 (Moustache sculpin) * '' Triglops nybelini'' A. S. Jensen, 1944 (Bigeye sculpin) * ''
Triglops pingelii ''Triglops'' is a genus of marine ray-finned fishes belonging to the Family (biology), family Cottidae, the typical sculpins. These fishes are found in the North Pacific, Arctic and North Atlantic Oceans. Taxonomy ''Triglops'' was first proposed ...
'' J. C. H. Reinhardt, 1837 (Ribbed sculpin) * '' Triglops scepticus'' Gilbert, 1896 (Spectacled sculpin) * '' Triglops xenostethus'' Gilbert, 1896 (Scaly-breasted sculpin)


Characteristics

''Triglops'' sculpins have slender, elongate, cylindrical to compressed or flattened bodies with a small head, which may also be slightly depressed or compressed, with a small horizontal or slightly oblique mouth. They have clearly separated
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...
s, the first containing between 9 and 13 slender spines and the second having between 19 and 31 soft rays. The
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
is supported by 18 to 32 soft rays. This genus is distinguished from the other marine sculpin genera by having the anus positioned halfway between the origins of the pelvic and anal fins, a feature shared with the genera ''
Clinocottus ''Clinocottus'' is a genus of sculpins, nearshore benthic fishes native to the northeastern Pacific Ocean. They are mentioned as sharpnose sculpins. Molecular data indicate that this genus is polyphyletic or paraphyletic, it does not constitute ...
'' and '' Leiocottus''. The
lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...
is made up of large scales which resemble plates and the scales underneath these form obvious rows of very small serrated plates, these are within close-set diagonal skin folds, a feature shared solely with ''
Jordania The longfin sculpin (''Jordania zonope'') is a species of marine ray-finned fish belonging to the family Jordaniidae, a small family of sculpins. This species is found in the northeastern Pacific Ocean. This species is the only known member of ...
''. There are four preopercular spines and the branchiostegal membranes are joined but do not connect to the isthmus. They have
vomerine teeth The vomer (; lat, vomer, lit=ploughshare) is one of the unpaired facial bones of the skull. It is located in the midsagittal line, and articulates with the sphenoid, the ethmoid, the left and right palatine bones, and the left and right maxill ...
but no palatine teeth. The
pelvic fin Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two en ...
has a single spine and three soft rays. There are no cutaneous cirri on the head. The fishes in this genus range from a maximum published
standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...
of in ''T. xenostethus'' up to in ''T. scepticus''.


Distribution and habitat

''Triglops'' sculpins are widespread in northern seas. In the Pacific Ocean they occur from the
Sea of Japan The Sea of Japan is the marginal sea between the Japanese archipelago, Sakhalin, the Korean Peninsula, and the mainland of the Russian Far East. The Japanese archipelago separates the sea from the Pacific Ocean. Like the Mediterranean Sea, it h ...
and
Sea of Okhotsk The Sea of Okhotsk ( rus, Охо́тское мо́ре, Ohótskoye móre ; ja, オホーツク海, Ohōtsuku-kai) is a marginal sea of the western Pacific Ocean. It is located between Russia's Kamchatka Peninsula on the east, the Kuril Islands ...
east into the
Bering Sea The Bering Sea (, ; rus, Бе́рингово мо́ре, r=Béringovo móre) is a marginal sea of the Northern Pacific Ocean. It forms, along with the Bering Strait, the divide between the two largest landmasses on Earth: Eurasia and The Ameri ...
, the
Aleutian Islands The Aleutian Islands (; ; ale, Unangam Tanangin,”Land of the Aleuts", possibly from Chukchi language, Chukchi ''aliat'', "island"), also called the Aleut Islands or Aleutic Islands and known before 1867 as the Catherine Archipelago, are a cha ...
the Gulf of Alaska south to
Puget Sound Puget Sound ( ) is a sound of the Pacific Northwest, an inlet of the Pacific Ocean, and part of the Salish Sea. It is located along the northwestern coast of the U.S. state of Washington. It is a complex estuarine system of interconnected ma ...
. In the Arctic they are found from the Chuckchi and
Beaufort Sea The Beaufort Sea (; french: Mer de Beaufort, Iñupiaq: ''Taġiuq'') is a marginal sea of the Arctic Ocean, located north of the Northwest Territories, the Yukon, and Alaska, and west of Canada's Arctic islands. The sea is named after Sir Fr ...
s east to the
Queen Elizabeth Islands The Queen Elizabeth Islands (french: Îles de la Reine-Élisabeth; formerly Parry Islands or Parry Archipelago) are the northernmost cluster of islands in Canada's Arctic Archipelago, split between Nunavut and the Northwest Territories in Northe ...
and
Hudson Bay Hudson Bay ( crj, text=ᐐᓂᐯᒄ, translit=Wînipekw; crl, text=ᐐᓂᐹᒄ, translit=Wînipâkw; iu, text=ᑲᖏᖅᓱᐊᓗᒃ ᐃᓗᐊ, translit=Kangiqsualuk ilua or iu, text=ᑕᓯᐅᔭᕐᔪᐊᖅ, translit=Tasiujarjuaq; french: b ...
and into the Atlantic south as far as Scotland and east to the
Barents Sea The Barents Sea ( , also ; no, Barentshavet, ; russian: Баренцево море, Barentsevo More) is a marginal sea of the Arctic Ocean, located off the northern coasts of Norway and Russia and divided between Norwegian and Russian territo ...
,
Kara Sea The Kara Sea (russian: Ка́рское мо́ре, ''Karskoye more'') is a marginal sea, separated from the Barents Sea to the west by the Kara Strait and Novaya Zemlya, and from the Laptev Sea to the east by the Severnaya Zemlya archipelago. ...
and Laptev Sea. They are benthic fishes which are found over varied substrates including rock, gravel, mud or sand at depths varying from the surface down to , although they are commonest between .


Biology

''Triglops'' sculpins are predators of invertebrates such as polychaetes,
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s and sometimes smaller fishes. They
spawn Spawn or spawning may refer to: * Spawn (biology), the eggs and sperm of aquatic animals Arts, entertainment, and media * Spawn (character), a fictional character in the comic series of the same name and in the associated franchise ** '' Spawn: Ar ...
from late summer into the winter, laying between 100 and over 2,500 demersal eggs.


References

{{Authority control Cottinae Taxa named by Johan Reinhardt Ray-finned fish genera Marine fish genera