Trichomanes Anomalum

''Trichomanes'' is a genus of ferns in the family Hymenophyllaceae, termed bristle ferns. The circumscription of the genus is disputed. All ferns in the genus are filmy ferns, with leaf tissue typically 2 cells thick. This thinness generally necessitates a permanently humid habitat, and makes the fronds somewhat translucent. Because of this membrane-like frond tissue, the plant is prone to drying out. “Filmy ferns” in the taxa Hymenophyllaceae grow in constantly wet environments. Many are found in cloud forests such as “Choco” in Colombia. There are also members of the taxa that can grow submersed in water.

The name bristle fern refers to the small bristle that protrudes from the indusia of these ferns.

Taxonomy

The genus ''Trichomanes'' was first described by Carl Linnaeus in 1753. Its status, like other genera in the subfamily Trichomanoideae, remains disputed. The Pteridophyte Phylogeny Group classification of 2016 (PPG I) accepts the genus as one of eight in the subfamily Trichomanoideae, saying that there are about 60 species. , the ''Checklist of Ferns and Lycophytes of the World'' listed about 70 species. Other sources, including ''Plants of the World Online'' , treated ''Trichomanes'' as the only genus in the subfamily Trichomanoideae, so that it included all the other genera, and had about 250 species.

Species

, the ''Checklist of Ferns and Lycophytes of the World'' accepted the following species:

*'' Trichomanes accedens'' C.Presl
*''Trichomanes alatum'' Sw.
*'' Trichomanes anadromum'' Rosenst.
*'' Trichomanes ankersii'' Parker
*'' Trichomanes anomalum'' Maxon & Morton
*''Trichomanes arbuscula'' Desv.
*''Trichomanes aureovestitum'' Proctor
*''Trichomanes bancroftii'' Hook. & Grev.
*''Trichomanes bicorne'' Hook.
*''Trichomanes bissei'' C.Sánchez
*''Trichomanes boivinii'' Bosch
*''Trichomanes botryoides'' Kaulf.
*''Trichomanes caliginum'' Lellinger
*''Trichomanes crassipilis'' Weath.
*''Trichomanes crinitum'' Sw.
*''Trichomanes crispiforme'' Alston
*'' Trichomanes crispum'' L.
*'' Trichomanes cristatum'' Kaulf.
*''Trichomanes cupressoides'' Desv.
*'' Trichomanes dactylites'' Sodiro
*'' Trichomanes delicatum'' Bosch
*'' Trichomanes diversifrons'' (Bory) Mett.
*'' Trichomanes ebiharae'' Fraser-Jenk. & Kholia
*'' Trichomanes egleri'' P.G.Windisch
*''Trichomanes elegans'' Rich.
*'' Trichomanes galeottii'' E.Fourn.
*''Trichomanes guidoi'' P.G.Windisch
*'' Trichomanes holopterum'' Kunze
*'' Trichomanes hostmannianum'' (Klotzsch) Kunze
*'' Trichomanes humboldtii'' (Bosch) Lellinger
*'' Trichomanes jenmanii'' Lellinger
*'' Trichomanes kalbreyeri'' Baker
*'' Trichomanes killipii'' Weath.
*'' Trichomanes laciniosum'' Alston
*''Trichomanes lozanoi'' M.T.Murillo
*'' Trichomanes lucens'' Sw.
*'' Trichomanes ludovicianum'' Rosenst.
*''Trichomanes macilentum'' Bosch
*''Trichomanes madagascariense'' (Bosch) Moore
*'' Trichomanes martiusii'' C.Presl
*'' Trichomanes micayense'' Hieron.
*''Trichomanes mougeotii'' Bosch
*''Trichomanes murilloanum'' A.Rojas
*'' Trichomanes osmundoides'' DC. ex Poir.
*'' Trichomanes padronii'' Proctor
*'' Trichomanes paucisorum'' R.C.Moran & B.Øllg.
*'' Trichomanes pedicellatum'' Desv.
*''Trichomanes pellucens'' Kunze
*'' Trichomanes pilosum'' Raddi
*'' Trichomanes pinnatifidum'' Bosch
*'' Trichomanes pinnatum'' Hedw.
*'' Trichomanes plumosum'' Kunze
*'' Trichomanes poeppigii'' C.Presl
*'' Trichomanes polypodioides'' L.
*'' Trichomanes procerum'' Fée
*''Trichomanes resinosum'' R.C.Moran
*''Trichomanes ribae'' Pacheco
*''Trichomanes robustum'' E.Fourn.
*'' Trichomanes roraimense'' Jenman
*'' Trichomanes scandens'' L.
*'' Trichomanes sellowianum'' C.Presl
*'' Trichomanes spruceanum'' Hook.
*'' Trichomanes steyermarkii'' P.G.Windisch & A.R.Sm.
*'' Trichomanes sublabiatum'' Bosch
*''Trichomanes superbum'' Bosch
*''Trichomanes tanaicum'' Hook.
*'' Trichomanes trichopodium'' A.Rojas
*''Trichomanes trigonum'' Desv.
*''Trichomanes trollii'' Bergdolt
*''Trichomanes tuerckheimii'' Christ
*''Trichomanes vandenboschii'' P.G.Windisch
*''Trichomanes vaupesensis'' Lellinger
*'' Trichomanes vittaria'' DC.

Species formerly placed in this genus include:

* ''Trichomanes angustatum'' Carmich. = ''Polyphlebium angustatum'' (Carmich.) Ebihara & Dubuisson
* ''Trichomanes bauerianum'' = '' Callistopteris baueriana'' (Endl.) Copel – Bauer's bristle fern
* ''Trichomanes boryanum'' = ''Cephalomanes atrovirens''
* ''Trichomanes boschianum'' = ''Vandenboschia boschiana'' (J.W.Sturm ex Bosch) Ebihara & K.Iwats. – Appalachian bristle fern
* ''Trichomanes caudatum'' Brack. = ''Abrodictyum caudatum'' (Brack.) Ebihara & K.Iwats.
* ''Trichomanes intricatum'' = '' Crepidomanes intricatum'' (Farrar) Ebihara & Weakley – Weft fern
* ''Trichomanes melanopus'' = '' Didymoglossum melanopus'' (Baker) Copel.
* ''Trichomanes petersii'' = '' Didymoglossum petersii'' (A. Gray) Copel. – Dwarf bristle fern
* ''Trichomanes reniforme'' = ''Hymenophyllum nephrophyllum'' Ebihara & K. Iwats.
* ''Trichomanes speciosum'' = ''Vandenboschia speciosa'' (Willd.) G. Kunkel
* ''Trichomanes tenuissimum'' = ''Polyphlebium tenuissimum'' (Bosch) ''comb. ined.''
* ''Trichomanes venosum'' R.Br. = ''Polyphlebium venosum'' (R.Br.) Copel.

References

{{Taxonbar, from=Q2704784

Hymenophyllales
Fern genera
Taxonomy articles created by Polbot