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Trachelidae is a family of araneomorph
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s (more recently evolved spiders with inward-pointing chelicerae) first described by
Eugène Simon Eugène Louis Simon (; 30 April 1848 – 17 November 1924) was a French naturalist who worked particularly on insects and spiders, but also on birds and plants. He is by far the most prolific spider taxonomist in history, describing over 4, ...
in 1897 as a subfamily called "Tracheleae" ("Trachelinae" in modern terminology). The Trachelidae family, also known as "ground sac spiders", is within the group of spiders known as the
RTA clade The RTA clade is a clade of araneomorph spiders, united by the possession of a retrolateral tibial apophysis – a backward-facing projection on the tibia of the male pedipalp. The clade contains over 21,000 species, almost half the current total ...
, which includes mostly wandering spiders that do not use webs. Spiders in the Trachelidae family are characterized as being 3-10mm long and having a red cephalothorax and a yellow/tan abdomen. They are commonly found indoors. It was placed in the family
Clubionidae The sac spiders of the family Clubionidae have a very confusing taxonomic history. Once, this family was a large catch-all taxon for a disparate collection of spiders, similar only in that they had eight eyes arranged in two rows and conical anter ...
, then later in Corinnidae when the Clubionidae were split up. The first study that suggested Trachelidae should be considered its own family was done by Deeleman-reinhold in 2001 as part of an analysis of RTA Clade spiders. An analysis by Martín J. Ramírez in 2014 suggested that it was not closely related to other members of the Corinnidae, and was better treated as a separate family. It was then placed in the CTC clade of spiders, or the Claw Tuft Clasper clade, which is a group of spiders that have two tarsal claws with tufts of hair. A major synapomorphy of Trachelidae is the reduction of leg spines. Other synapomorphies of the family include no scales, no epiandrous spigot, only one major ampullate gland in females, no median apophysis, and the secondary spermatheca are the same size as the primary. Currently, there are 271 known species across 20 genera.


Genera

, the
World Spider Catalog The World Spider Catalog (WSC) is an online searchable database concerned with spider taxonomy. It aims to list all accepted families, genera and species, as well as provide access to the related taxonomic literature. The WSC began as a series of ...
accepts the following genera: *'' Afroceto'' Lyle & Haddad, 2010 – Africa *''
Capobula ''Capobula'' is a genus of southern African spiders in the family Trachelidae. The type species was first described by Eugène Simon from a juvenile found in South Africa. It was placed with '' Orthobula'' for several morphological similarities, ...
'' Haddad, Jin, Platnick & Booysen, 2021 – South Africa, Lesotho *''
Cetonana ''Cetonana'' is a genus of araneomorph spiders in the family Trachelidae, first described by Embrik Strand in 1929. Species it contains five species: *''Cetonana laticeps ''Cetonana'' is a genus of araneomorph spiders in the family Trachelid ...
'' Strand, 1929 – Russia, Brazil *'' Fuchiba'' Haddad & Lyle, 2008 – Africa *''
Fuchibotulus ''Fuchibotulus'' is a genus of South African and Mozambican araneomorph spiders in the family Trachelidae Trachelidae is a family of araneomorph spiders (more recently evolved spiders with inward-pointing chelicerae) first described by Eugè ...
'' Haddad & Lyle, 2008 – South Africa, Mozambique *'' Jocquestus'' Lyle & Haddad, 2018 – Africa *''
Meriola ''Meriola'' is a genus of araneomorph spiders in the family Trachelidae, first described by Nathan Banks in 1895. Species it contains twenty-four species, three from the United States and Canada: *'' Meriola arcifera'' (Simon, 1886) – Chile, ...
'' Banks, 1895 – South America, North America, Guatemala *'' Metatrachelas'' Bosselaers & Bosmans, 2010 – Algeria, Europe *''
Orthobula ''Orthobula'' is a genus of araneomorph spiders first described by Eugène Simon in 1897 as a member of Liocranidae. It was transferred to Corinnidae in 2002, to Phrurolithidae in 2014, and to Trachelidae in 2017. Species it contains twenty sp ...
'' Simon, 1897 – Asia, Africa *''
Paccius ''Paccius'' is a genus of African araneomorph spiders first described by Eugène Simon in 1898 as a member of Corinnidae, and moved to Trachelidae in 2014. Species it contains eight species from Madagascar and Seychelles Seychelles (, ...
'' Simon, 1898 – Madagascar, Seychelles *''
Paraceto ''Paraceto'' is a genus of Chinese and Korean araneomorph spiders in the family Trachelidae Trachelidae is a family of araneomorph spiders (more recently evolved spiders with inward-pointing chelicerae) first described by Eugène Simon in 189 ...
'' Jin, Yin & Zhang, 2017 – China, Korea *''
Paratrachelas ''Paratrachelas'' is a genus of araneomorph spiders in the family Trachelidae Trachelidae is a family of araneomorph spiders (more recently evolved spiders with inward-pointing chelicerae) first described by Eugène Simon in 1897 as a subfami ...
'' Kovblyuk & Nadolny, 2009 – Asia, Algeria, Europe *''
Patelloceto ''Patelloceto'' is a genus of African araneomorph spiders in the family Trachelidae Trachelidae is a family of araneomorph spiders (more recently evolved spiders with inward-pointing chelicerae) first described by Eugène Simon in 1897 as a s ...
'' Lyle & Haddad, 2010 – Ethiopia *''
Planochelas ''Planochelas'' is a genus of African araneomorph spiders in the family Trachelidae, first described by R. Lyle & C. R. Haddad in 2009. it contains only three species from Ghana, Uganda }), is a landlocked country in East Africa. The co ...
'' Lyle & Haddad, 2009 – Ghana, Uganda, Ivory Coast *''
Poachelas ''Poachelas'' is a genus of African araneomorph spiders in the family Trachelidae Trachelidae is a family of araneomorph spiders (more recently evolved spiders with inward-pointing chelicerae) first described by Eugène Simon in 1897 as a su ...
'' Haddad & Lyle, 2008 – South Africa, Zimbabwe *''
Spinotrachelas ''Spinotrachelas'' is a genus of South African araneomorph spiders in the family Trachelidae, first described by C. R. Haddad in 2006. Species it contains five species, all found in South Africa South Africa, officially the Republic of ...
'' Haddad, 2006 – South Africa *''
Thysanina ''Thysanina'' is a genus of African araneomorph spiders first described by Eugène Simon in 1910. Originally placed with the Liocranidae, it was moved to the Corinnidae in 2000, and to the Trachelidae Trachelidae is a family of araneomorph sp ...
'' Simon, 1910 – South Africa, Namibia, Tanzania *'' Trachelas'' L. Koch, 1872 – Asia, Central America, North America, Caribbean, Spain, South America, Africa *'' Trachelopachys'' Simon, 1897 – South America *'' Utivarachna'' Kishida, 1940 – Asia


References

Araneomorphae families {{Trachelidae-stub