The tourmaline sunangel (''Heliangelus exortis'') is a species of

hummingbird Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics aro ...

in the "coquettes", tribe

Lesbiini Lesbiini is one of the two tribes that make up the subfamily Lesbiinae in the hummingbird family Trochilidae. The other tribe is Heliantheini (brilliants). The informal name "coquettes" has been proposed for this group as the largest genus, ''L ...

of subfamily Lesbiinae. It is found in

Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...

and

Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...

.HBW and BirdLife International (2020) ''Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world'' Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip xls zipped 1 MBretrieved May 27, 2021

Taxonomy and systematics

The tourmaline sunangel is

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...

. However, at times it has been considered conspecific with the

flame-throated sunangel The flame-throated sunangel or little sunangel (''Heliangelus micraster'') is a species of hummingbird in the "coquettes", tribe Lesbiini of subfamily Lesbiinae. It is found in Ecuador and Peru.HBW and BirdLife International (2020) ''Handbook o ...

(''H. micraster'').Heynen, I. and P. F. D. Boesman (2020). Tourmaline Sunangel (''Heliangelus exortis''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.tousun1.01 retrieved January 21, 2022

Description

The tourmaline sunangel is long. Males weigh and females . Their bill is blackish, straight, and of medium length. Both sexes have dark shining green upperparts and dusky gray underparts with round green spots. Their tails' central feathers are dark bronze and the others blackish. Adult males have a glittering green frontlet just above the bill, a dark purple-blue chin, and a glittering rosy pink

gorget A gorget , from the French ' meaning throat, was a band of linen wrapped around a woman's neck and head in the medieval period or the lower part of a simple chaperon hood. The term later described a steel or leather collar to protect the thro ...

with a glittering emerald green border below it. Adult females lack the green frontlet, their chin is blackish, and their throat whitish with green to dusky speckles and sometimes a few glittering rosy feathers. Juveniles are like the adult female but the male has a white chin.

Distribution and habitat

The tourmaline sunangel is found in all three

Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...

ranges of Colombia and south on the eastern slope to Ecuador's Morona-Santiago Province. It inhabits the interior and edges of humid to wet premontane forest, especially

cloudforest A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest (TMCF), is a generally tropical or subtropical, evergreen, montane, moist forest characterized by a persistent, frequent or seasonal low-level cloud c ...

and

elfin forest Dwarf forest, elfin forest, or pygmy forest is an uncommon ecosystem featuring miniature trees, inhabited by small species of fauna such as rodents and lizards. They are usually located at high elevations, under conditions of sufficient air humi ...

. It also occurs in bushy clearings and shrubby pastures. In elevation it mainly ranges between but is found as low as .

Behavior

Movement

The tourmaline sunangel is almost entirely sendentary but in Colombia makes occasional movements into the tropical zone.

Feeding

The tourmaline sunangel is aggressively territorial at feeding locations. It takes nectar from a variety of flowering plants, usually those low to the ground both in semi-open landscapes and within the forest. It usually clings to flowers to feed rather than hovering at them. It captures insects by

hawking Hawking may refer to: People * Stephen Hawking (1942–2018), English theoretical physicist and cosmologist *Hawking (surname), a family name (including a list of other persons with the name) Film * ''Hawking'' (2004 film), about Stephen Haw ...

from a perch and by gleaning from vegetation.

Breeding

The tourmaline sunangel's breeding season appears to span from March to August. The clutch of two white eggs is incubated by the female. Nothing else is known about the species' breeding

phenology Phenology is the study of periodic events in biological life cycles and how these are influenced by seasonality, seasonal and interannual variations in climate, as well as environmental factor, habitat factors (such as elevation). Examples includ ...

Vocalization

A vocalization believed to be the tourmaline sunangel's song is "a repeated series of single notes...typically a few high-pitched upslurred notes followed by a slightly lower-pitched downslurred note, 'suwee....suwee..tsew...suwee...tsew...'." Its call is "a short dry gravelly trill 'trrr', repeated at intervals" and it also makes a continuous dry chatter.

Status

The

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

has assessed the tourmaline sunangel as being of Least Concern. Its population size is not known but is believed to be stable. It is locally fairly common and occurs in several protected areas.

References

{{Taxonbar, from=Q1263086

tourmaline sunangel The tourmaline sunangel (''Heliangelus exortis'') is a species of hummingbird in the "coquettes", tribe Lesbiini of subfamily Lesbiinae. It is found in Colombia and Ecuador.HBW and BirdLife International (2020) ''Handbook of the Birds of the Wo ...

Birds of the Colombian Andes Birds of the Ecuadorian Andes

Taxonomy articles created by Polbot