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The toolache wallaby or Grey's wallaby (''Notamacropus greyi'') is an extinct species of
wallaby A wallaby () is a small or middle-sized macropod native to Australia and New Guinea, with introduced populations in New Zealand, Hawaii, the United Kingdom and other countries. They belong to the same taxonomic family as kangaroos and som ...
from southeastern
South Australia South Australia (commonly abbreviated as SA) is a state in the southern central part of Australia. It covers some of the most arid parts of the country. With a total land area of , it is the fourth-largest of Australia's states and territories ...
and southwestern
Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...
.


Taxonomy

A species described by George Waterhouse in 1846. The type specimen was collected at Coorong in South Australia. The author cites an earlier name, ''Halmaturus greyii'', published by
John Edward Gray John Edward Gray, FRS (12 February 1800 – 7 March 1875) was a British zoologist. He was the elder brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray (1766–1828). The same is used for ...
in 1843 without a valid description, assigning it to a subgenus of the same name—''Macropus'' (''Halmaturus'')—and providing the common name of the newly described species as Grey's wallaby. The common name and epithet ''greyi'' commemorates the collector and explorer
George Grey Sir George Grey, KCB (14 April 1812 – 19 September 1898) was a British soldier, explorer, colonial administrator and writer. He served in a succession of governing positions: Governor of South Australia, twice Governor of New Zealand, Go ...
, who provided the two specimens to researchers at the
British Museum of Natural History The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum ...
. A systematic revision has seen the species placed in a subgeneric arrangement as ''Macropus'' (''Notamacropus'') ''greyi'', recognising an affinity with eight other species of the subgenus named as ''
Notamacropus ''Notamacropus'' is a genus of small marsupials in the family Macropodidae, commonly known as wallabies (among other species). In 2019, a reassessment of macropod taxonomy determined that ''Notamacropus'' and ''Osphranter'', formerly considered ...
'' Dawson and Flannery, 1985. An arrangement that elevates the subgenera of ''Macropus'' is recognised as ''Notamacropus greyi''. A genetic analysis found that its closest relative is the
western brush wallaby The western brush wallaby (''Notamacropus irma''), also known as the black-gloved wallaby, is a species of wallaby found in the southwestern coastal region of Western Australia. The wallaby's main threat is predation by the introduced red fox (' ...
. The common names have also included monkeyface and onetwo.


Description

The toolache wallaby was a slim, graceful, and elegant creature that had a pale ashy-brown pelt with a buff-yellow underbelly. The tail was pale grey and became almost white near the tip. The distinct black mark on its face reached from its nose to the eye. The forearms, feet, and tips of the ears were also black. The different colours of the animal also consisted of different textured furs which are believed to have changed seasonally or varied depending on the individual. The body measurements differed between males and females. In general, male toolache wallabies had a head and body length up to 810 mm while females measured up as 840 mm. Despite the females being taller, males had longer tail lengths at about 730 mm while the female's tail length was 710 mm.


Behaviour

The toolache wallaby was a nocturnal animal, foraging for vegetation during the twilight hours of the day. Their movements were unusual and extremely rapid, able to outpace almost any terrestrial predator; they were known to evade the fastest dogs of the colonial hunters.


Habitat

The toolache wallaby occupied the southeastern corner of Australia to the western part of Victoria. The preferred habitat ranged from swampy short grassland areas, to taller grassed areas of the open country. Toolache wallabies were known to be sociable creatures who lived in groups; they were often seen resting and grazing in groups.


Threats

A combination of numerous threats caused the decline and eventual extinction of the toolache wallaby. One of the largest factors was the destruction of its habitat. Since swamps were an important part of its habitat, once they were cleared out, much of the vegetation went with it. Besides the destruction of its habitat, the introduction of predators, such as the European red fox, began to kill off the species as well. Additionally, the animal was also hunted for sport and for its pelt. A lesser known fact about extinction threats to the species was the effect of the direct predation from
Aboriginal Australians Aboriginal Australians are the various Indigenous peoples of the Australian mainland and many of its islands, such as Tasmania, Fraser Island, Hinchinbrook Island, the Tiwi Islands, and Groote Eylandt, but excluding the Torres Strait Isl ...
and their prehistoric introduction of dingos where macropods generally resided.


Extinction

The toolache wallaby only survived 85 years after European occupation. In the 1920s, a conservation effort was made to try and bring the animal back from the brink of extinction. The plan was to capture and breed the last known surviving members of the species in captivity. This effort ended in disaster after 10 of the 14 of them were accidentally killed in the attempt to capture them. The remaining four survived in captivity. The last wild sightings were recorded in 1924, and the last known toolache wallaby survived in captivity until 1939. The species is presumed to be extinct, although extensive research is still being conducted in the region after reports of suspected sightings through the 1970s. However, no members of the species have been sighted since.


References

* {{Taxonbar, from=Q109262320, from2=Q209153 Macropods Extinct marsupials Mammal extinctions since 1500 Extinct mammals of South Australia Mammals of Victoria (Australia) Marsupials of Australia Mammals described in 1846 Species made extinct by human activities