Tomoceridae
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The Entomobryomorpha are one of the three main groups (
order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...
) of
springtail Springtails (Collembola) form the largest of the three lineages of modern hexapods that are no longer considered insects (the other two are the Protura and Diplura). Although the three orders are sometimes grouped together in a class called Ento ...
s (Collembola), tiny hexapods related to
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s. This group was formerly treated as a
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
, the Entomobryoidea. They can be best distinguished from the other springtail groups by their body shape. The
Symphypleona The order Symphypleona is one of the three main groups of springtails (Collembola), tiny hexapods related to insects. When the springtails were still believed to be an order of insects, the Symphypleona were ranked as a suborder. They can be ...
are very round animals, almost
spherical A sphere () is a geometrical object that is a three-dimensional analogue to a two-dimensional circle. A sphere is the set of points that are all at the same distance from a given point in three-dimensional space.. That given point is the ce ...
. The
Poduromorpha The order Poduromorpha is one of the three main groups of springtails (Collembola), tiny hexapods related to insects. This group was formerly treated as a superfamily Poduroidea. They can be best distinguished from the other springtail groups ...
are also very plump but have a more oval shape. The Entomobryomorpha, by contrast, contain the slimmest springtails. They either have short legs and
antenna Antenna ( antennas or antennae) may refer to: Science and engineering * Antenna (radio), also known as an aerial, a transducer designed to transmit or receive electromagnetic (e.g., TV or radio) waves * Antennae Galaxies, the name of two collid ...
e, but their long bodies set them apart, or long legs and antennae, as well as well-developed
furcula The (Latin for "little fork") or wishbone is a forked bone found in most birds and some species of non-avian dinosaurs, and is formed by the fusion of the two pink clavicles. In birds, its primary function is in the strengthening of the thoracic ...
e; these are the most characteristic members of the order.


Systematics

The Entomobryomorpha were, as Entomobryoidea, united with the
Poduromorpha The order Poduromorpha is one of the three main groups of springtails (Collembola), tiny hexapods related to insects. This group was formerly treated as a superfamily Poduroidea. They can be best distinguished from the other springtail groups ...
(then called Poduroidea) in a group called "Arthropleona", but this has more recently turned out to be
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
. Actually, the Entomobryomorpha, the Poduromorpha, and the third springtaill lineage – the
Symphypleona The order Symphypleona is one of the three main groups of springtails (Collembola), tiny hexapods related to insects. When the springtails were still believed to be an order of insects, the Symphypleona were ranked as a suborder. They can be ...
– are equally distinct from each other. Their treatment at equal
taxonomic rank In biological classification, taxonomic rank is the relative level of a group of organisms (a taxon) in an ancestral or hereditary hierarchy. A common system consists of species, genus, family (biology), family, order (biology), order, class (b ...
reflects this. Their rank has also varied a bit. When the springtails were still believed to be an order of insects, the "Arthropleona" and the Symphypleona were treated as
suborder Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...
s.


List of families

The superfamilies and
families Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Ideall ...
are arranged in the presumed
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
ary sequence. The list presented here follows a 2008 review of the Entomobryomorpha. The review abolishes the former "Actaletoidea", which is apparently
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
; its namesake family is now placed in the Isotomoidea, while the Coenaletidae form a new
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
superfamily. The former Cyphoderidae are demoted to a
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
of the
Paronellidae Paronellidae is a family of elongate-bodied springtails in the order Entomobryomorpha. There are about 18 genera and at least 90 described species in Paronellidae. Genera These 18 genera belong to the family Paronellidae: * '' Callyntrura'' Bö ...
. Superfamily Isotomoidea * Family Isotomidae * Family Actaletidae * Family Protentomobryidae (
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
) Superfamily Coenaletoidea * Family Coenaletidae Superfamily Tomoceroidea * Family Oncopoduridae * Family Tomoceridae Superfamily
Entomobryoidea :''Entomobryoidea is also the old name of Entomobryomorpha, when these were placed in the "Arthropleona" The Entomobryoidea are a superfamily of springtails (Collembola), tiny hexapods related to insects. In the modern sense, this group is placed ...
* Family
Paronellidae Paronellidae is a family of elongate-bodied springtails in the order Entomobryomorpha. There are about 18 genera and at least 90 described species in Paronellidae. Genera These 18 genera belong to the family Paronellidae: * '' Callyntrura'' Bö ...
– includes Cyphoderidae * Family
Entomobryidae Entomobryidae, sometimes called "slender springtails", is a family of springtails characterised by having an enlarged fourth abdominal segmentation (biology), segment and a well-developed furcula (Collembola), furcula. Species in this family ma ...
* Family
Microfalculidae :''Entomobryoidea is also the old name of Entomobryomorpha, when these were placed in the "Arthropleona" The Entomobryoidea are a superfamily of springtails (Collembola), tiny hexapods related to insects. In the modern sense, this group is place ...
* Family Oncobryidae (fossil) * Family
Praentomobryidae :''Entomobryoidea is also the old name of Entomobryomorpha, when these were placed in the "Arthropleona" The Entomobryoidea are a superfamily of springtails (Collembola), tiny hexapods related to insects. In the modern sense, this group is placed ...
(fossil)


See also

*''
Tomocerus minor ''Tomocerus minor'' is a species of springtail in the family Tomoceridae. It is widespread from the Arctic to Europe, Western Asia, Western and Central Asia down to the Sino-Japanese, Northern and Pacific North America, Hawaii, the Caribbean mai ...
''


References

{{Taxonbar, from=Q1045208 Arthropod orders