''Tomarus'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...

, and a few species occur in the

Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...

.Ratcliffe, B. C. and R. D. Cave. (2010)
The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of ''Tomarus adoceteus'', new species (Pentodontini) and ''Caymania nitidissima'', new genus and species (Phileurini).
''Insecta Mundi'' 139, 1-15. The adult beetles are nocturnal and attracted to lights; the

larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...

e generally remain in the soil, often feeding on plant roots. In order to identify species, the

paramere Parameres ('side parts') are part of the external reproductive organs of male insects and the term was first used by Karl Wilhelm Verhoeff, Verhoeff in 1893 for the lateral genital lobes in Coleoptera. The primary phallic lobes which appear in th ...

s of the male must be pulled out of the abdomen and examined. This process should be done carefully, because the parameres are quite fragile, "almost

parchment Parchment is a writing material made from specially prepared untanned skins of animals—primarily sheep, calves, and goats. It has been used as a writing medium for over two millennia. Vellum is a finer quality parchment made from the skins of ...

-like".

Species

The following species belong to the genus ''Tomarus'': * '' Tomarus adoceteus'' Ratcliffe & Cave, 2010^{c g} * '' Tomarus bidentulus'' (Fairmaire, 1892)^c * '' Tomarus bituberculatus'' (Palisot De Beauvois, 1805)^{c g} * '' Tomarus burmeisteri'' (Steinheil, 1872)^{c g} * '' Tomarus cicatricosus'' (Prell, 1937)^{c g} * '' Tomarus colombianus'' Lopez-Garcia & Gasca-Alvarez, 2014^g * ''

Tomarus columbianus ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...

'' López-García & Gasca-Álvarez, 2014^{c g} * '' Tomarus compositus'' (Wickham, 1911)^{c g} * '' Tomarus cuniculus'' (Fabricius, 1801)^{i c g b} * '' Tomarus discrepans'' Escalona & Joly, 2006^{c g} * '' Tomarus ebenus'' (Degeer, 1774)^{c g} = type species * '' Tomarus effetus'' (Wickham, 1914)^{c g} * '' Tomarus formosianus'' Grouvelle, 1913^g * '' Tomarus fossor'' (Latreille, 1833)^{c g} * ''

Tomarus gyas ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...

'' Erichson, 1848^{c g} * ''

Tomarus laevicollis ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...

'' (Bates, 1888)^{i c g} * ''

Tomarus maimon ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...

'' Erichson, 1847^{c g} * ''

Tomarus maternus ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...

'' (Prell, 1937)^{c g} * ''

Tomarus nasutus ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...

'' (Burmeister, 1847)^{i c g} * ''

Tomarus neglectus ''Tomarus neglectus'' is a species of rhinoceros beetle in the family Scarabaeidae. References Further reading * Dynastinae Articles created by Qbugbot Beetles described in 1847 {{Dynastinae-stub ...

'' (LeConte, 1847)^{i c g b} * ''

Tomarus peruvianus ''Tomarus'' is a genus of scarabaeidae, scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the C ...

'' ( Endrödi, 1970)^c * '' Tomarus pullus'' (Prell, 1937)^{c g} * '' Tomarus pumilus'' (Prell, 1937)^{c g} * '' Tomarus roigjunenti'' Neita & Ratcliffe, 2017^{c g} * '' Tomarus rosettae'' ( Endrödi, 1968)^{c g} * '' Tomarus rostratus'' Dupuis, 2014^{c g} * '' Tomarus rubripes'' (Boheman, 1858)^{c g} * '' Tomarus sallaei'' (Bates, 1888)^{i c g b} * '' Tomarus selanderi'' (Cartwright, 1959)^{i c g} * '' Tomarus similis'' (Endrödi, 1968)^{c g} * '' Tomarus spinipenis'' Neita & Ratcliffe, 2017^{c g} * '' Tomarus subtropicus'' (Blatchley, 1922)^{i c g b} (sugarcane grub)Buss, E. A
Sugarcane Grub, ''Tomarus subtropicus'' Blatchley (Insecta: Coleoptera: Scarabaeidae).
EENY-318 (IN593). Entomology and Nematology. Florida Cooperative Extension Service. University of Florida IFAS. Published 2003, revised 2006. * '' Tomarus villosus'' (Burmeister, 1847)^{c g} Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net ;Note - now placed in the restored genus '' Ligyrus'': * '' Tomarus gibbosus'' (LeConte, 1856), the carrot beetle * '' Tomarus relictus'' (Say, 1825)

References

{{Taxonbar, from=Q15635796 Dynastinae Beetles of South America