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''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
.Ratcliffe, B. C. and R. D. Cave. (2010)
The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of ''Tomarus adoceteus'', new species (Pentodontini) and ''Caymania nitidissima'', new genus and species (Phileurini).
''Insecta Mundi'' 139, 1-15.
The adult beetles are nocturnal and attracted to lights; the larvae generally remain in the soil, often feeding on plant roots. In order to identify species, the
paramere Parameres ('side parts') are part of the external reproductive organs of male insects and the term was first used by Karl Wilhelm Verhoeff, Verhoeff in 1893 for the lateral genital lobes in Coleoptera. The primary phallic lobes which appear in th ...
s of the male must be pulled out of the abdomen and examined. This process should be done carefully, because the parameres are quite fragile, "almost parchment-like".


Species

The following species belong to the genus ''Tomarus'': * '' Tomarus adoceteus'' Ratcliffe & Cave, 2010 c g * '' Tomarus bidentulus'' (Fairmaire, 1892) c * '' Tomarus bituberculatus'' (Palisot De Beauvois, 1805) c g * '' Tomarus burmeisteri'' (Steinheil, 1872) c g * '' Tomarus cicatricosus'' (Prell, 1937) c g * ''
Tomarus colombianus ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean. ...
'' Lopez-Garcia & Gasca-Alvarez, 2014 g * ''
Tomarus columbianus ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...
'' López-García & Gasca-Álvarez, 2014 c g * '' Tomarus compositus'' (Wickham, 1911) c g * '' Tomarus cuniculus'' (Fabricius, 1801) i c g b * '' Tomarus discrepans'' Escalona & Joly, 2006 c g * ''
Tomarus ebenus ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...
'' (Degeer, 1774) c g = type species * ''
Tomarus effetus ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...
'' (Wickham, 1914) c g * ''
Tomarus formosianus ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...
'' Grouvelle, 1913 g * ''
Tomarus fossor ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...
'' (Latreille, 1833) c g * ''
Tomarus gyas ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...
'' Erichson, 1848 c g * ''
Tomarus laevicollis ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...
'' (Bates, 1888) i c g * ''
Tomarus maimon ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...
'' Erichson, 1847 c g * ''
Tomarus maternus ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...
'' (Prell, 1937) c g * ''
Tomarus nasutus ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...
'' (Burmeister, 1847) i c g * ''
Tomarus neglectus ''Tomarus neglectus'' is a species of rhinoceros beetle in the family Scarabaeidae. References Further reading * Dynastinae Articles created by Qbugbot Beetles described in 1847 {{Dynastinae-stub ...
'' (LeConte, 1847) i c g b * ''
Tomarus peruvianus ''Tomarus'' is a genus of scarabaeidae, scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the C ...
'' ( Endrödi, 1970) c * ''
Tomarus pullus ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...
'' (Prell, 1937) c g * '' Tomarus pumilus'' (Prell, 1937) c g * ''
Tomarus roigjunenti ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean. ...
'' Neita & Ratcliffe, 2017 c g * '' Tomarus rosettae'' ( Endrödi, 1968) c g * '' Tomarus rostratus'' Dupuis, 2014 c g * '' Tomarus rubripes'' (Boheman, 1858) c g * '' Tomarus sallaei'' (Bates, 1888) i c g b * '' Tomarus selanderi'' (Cartwright, 1959) i c g * ''
Tomarus similis ''Tomarus'' is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.Rat ...
'' (Endrödi, 1968) c g * '' Tomarus spinipenis'' Neita & Ratcliffe, 2017 c g * '' Tomarus subtropicus'' (Blatchley, 1922) i c g b (sugarcane grub)Buss, E. A
Sugarcane Grub, ''Tomarus subtropicus'' Blatchley (Insecta: Coleoptera: Scarabaeidae).
EENY-318 (IN593). Entomology and Nematology. Florida Cooperative Extension Service. University of Florida IFAS. Published 2003, revised 2006.
* '' Tomarus villosus'' (Burmeister, 1847) c g Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net ;Note - now placed in the restored genus '' Ligyrus'': * '' Tomarus gibbosus'' (LeConte, 1856), the carrot beetle * '' Tomarus relictus'' (Say, 1825)


References

{{Taxonbar, from=Q15635796 Dynastinae Beetles of South America