Tobago Main Ridge Forest Reserve
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Main Ridge is the main mountainous ridge on the island of
Tobago Tobago () is an List of islands of Trinidad and Tobago, island and Regions and municipalities of Trinidad and Tobago, ward within the Trinidad and Tobago, Republic of Trinidad and Tobago. It is located northeast of the larger island of Trini ...
,
Trinidad and Tobago Trinidad and Tobago (, ), officially the Republic of Trinidad and Tobago, is the southernmost island country in the Caribbean. Consisting of the main islands Trinidad and Tobago, and numerous much smaller islands, it is situated south of ...
. It is a chain of hills which runs from southwest to northeast between the
Caribbean Sea The Caribbean Sea ( es, Mar Caribe; french: Mer des Caraïbes; ht, Lanmè Karayib; jam, Kiaribiyan Sii; nl, Caraïbische Zee; pap, Laman Karibe) is a sea of the Atlantic Ocean in the tropics of the Western Hemisphere. It is bounded by Mexico ...
and the Southern Tobago fault system and reaches a maximum height of . The Main Ridge Forest Reserve, which was legally established in 1776, is one of the oldest
protected area Protected areas or conservation areas are locations which receive protection because of their recognized natural, ecological or cultural values. There are several kinds of protected areas, which vary by level of protection depending on the ena ...
s in the world. It is a popular site for birdwatching and
ecotourism Ecotourism is a form of tourism involving responsible travel (using sustainable transport) to natural areas, conserving the environment, and improving the well-being of the local people. Its purpose may be to educate the traveler, to provide funds ...
. Main Ridge provides important habitat for native plants and animals, including several species
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to Tobago.


History

The
Treaty of Paris Treaty of Paris may refer to one of many treaties signed in Paris, France: Treaties 1200s and 1300s * Treaty of Paris (1229), which ended the Albigensian Crusade * Treaty of Paris (1259), between Henry III of England and Louis IX of France * Trea ...
in 1763 ended Tobago's status as a neutral territory and made it a British colony. The Treaty of Aix-la-Chappelle in 1748 had designated Tobago neutral territory and left it in the hands of its remaining indigenous population, but the return to British control led to a rapid conversion of the island to a plantation economy. Under the direction of the
Board of Trade The Board of Trade is a British government body concerned with commerce and industry, currently within the Department for International Trade. Its full title is The Lords of the Committee of the Privy Council appointed for the consideration of ...
, the island was surveyed, divided into plots, and sold to planters. The upper portions of the Main Ridge were reserved as "Woods for the Protection of the Rains" and thus remained uncleared and uncultivated. The decision to preserve forests to maintain rainfall was driven by the efforts of
Soame Jenyns Soame Jenyns (1 January 1704 – 18 December 1787) was an English writer and Member of Parliament. He was an early advocate of the ethical consideration of animals. Life and work He was the eldest son of Sir Roger Jenyns and his second wife El ...
, a commissioner of the Board of Trade and Member of Parliament. Jenyns was convinced of the importance of forests for preserving rainfall through the work of Stephen Hales on
plant physiology Plant physiology is a subdiscipline of botany concerned with the functioning, or physiology, of plants. Closely related fields include plant morphology (structure of plants), plant ecology (interactions with the environment), phytochemistry (bi ...
and
transpiration Transpiration is the process of water movement through a plant and its evaporation from aerial parts, such as leaves, stems and flowers. Water is necessary for plants but only a small amount of water taken up by the roots is used for growth a ...
. It took Jenyns eleven years to convince the British Parliament of the importance of the endeavour, but on 13 April 1776, Parliament passed an ordinance establishing the reserve "for the purpose of attracting frequent showers of rain upon which the fertility of lands in these climates doth entirely depend". This action produced one of the oldest protected areas in the world geared towards conservation and has been described as "the first act in the modern environmental movement". Beginning in 1904, the rain reserve was combined with adjacent
Crown land Crown land (sometimes spelled crownland), also known as royal domain, is a territorial area belonging to the monarch, who personifies the Crown. It is the equivalent of an entailed estate and passes with the monarchy, being inseparable from it. ...
s and proclaimed the Main Ridge Forest Reserve.


Geography

Main Ridge forms the mountainous backbone of the island of Tobago, which is the smaller and more northern of the two main islands that make up the southern Caribbean nation of Trinidad and Tobago.. The chain of hills is long and runs from the southwest to the northeast, roughly parallel to the orientation of the island. Main Ridge reaches an elevation of above sea level at Centre Hill, but lacks any well-defined peaks. To the northwest it is bounded by the Caribbean Sea and by the Southern Tobago fault system to the southeast. Rainfall averages about per year. Most streams run along oblique-slip faults with a northwest orientation. Major streams draining the Main Ridge include the Coffee, Hillsborough, Goldsborough, and Queens Rivers. Streams which drain to the northwest are short and steep, while those which drain to the southeast are longer, with better-developed courses. Waterfalls, including the Argylle Waterfall (Tobago's highest) occur at major transitions between rock types. The northeastern slopes are steeper than the southwestern ones and are among the most landslide-prone parts of the island.


Geology

Main Ridge is primarily underlain by the North Coast Schist Group (NCSG), which occupies the northern third of the island of Tobago.. The island of Tobago is the main exposed portion of the Tobago terrane, a fragment of crustal material lying between the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
and
South American Plate The South American Plate is a major tectonic plate which includes the continent of South America as well as a sizable region of the Atlantic Ocean seabed extending eastward to the African Plate, with which it forms the southern part of the Mid-A ...
s. The rocks of the NCSG are
metavolcanic Metavolcanic rock is volcanic rock that shows signs of having experienced metamorphism. In other words, the rock was originally produced by a volcano, either as lava or tephra. The rock was then subjected to high pressure, high temperature or both ...
; the underlying igneous rock was laid down during the Late Jurassic or
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
. They underwent metamorphosis prior to the mid
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
, when additional volcanic intrusion formed the rocks of the Tobago Volcanic Group. A combination of uplift, erosion, and faulting during the late
Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceo ...
and
Paleogene The Paleogene ( ; British English, also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period, geologic period and system that spans 43 million years from the end of the Cretaceous Period million yea ...
led to the elevated horst block separated from the half
graben In geology, a graben () is a depressed block of the crust of a planet or moon, bordered by parallel normal faults. Etymology ''Graben'' is a loan word from German, meaning 'ditch' or 'trench'. The word was first used in the geologic contex ...
of the southern lowlands by the Southern Tobago fault system.


Vegetation and ecology

Main Ridge is dominated by lower montane rain forest (according to
John Stanley Beard John Stanley Beard (15 February 1916 – 17 February 2011) was a British-born forester and ecologist who resided in Australia. Beard studied at the University of Oxford where he completed his doctoral thesis on tropical forestry. While working ...
's classification of the vegetation of Tobago). In Tobago, these forests are characterised by an emergent canopy of '' Licania biglandulosa'' and ''
Byrsonima spicata ''Byrsonima'' is one of about 75 genera in the Malpighiaceae, a family (biology), family of flowering plants in the order Malpighiales. In particular in American English, they are known as locustberries (which also may refer specifically to ''Brs ...
'', with the palm ''
Euterpe broadwayi ''Euterpe broadwayi'', the manac, or manicol, is a tall, slender-stemmed, pinnate-leaved palm native to Trinidad and Tobago, Grenada, Saint Vincent and the Grenadines and Dominica. Stems usually grow in a cluster and are 8–20 metres tall and ...
'' more common on exposed ridges and summits. Moist lowland forest dominates the lower-lying portions of the Main Ridge. The forests experienced limited human disturbance prior to the 1940s, but were badly damaged by
Hurricane Flora Hurricane Flora is among the deadliest Atlantic hurricanes in recorded history, with a death total of at least 7,193. The seventh tropical storm and sixth hurricane of the 1963 Atlantic hurricane season, Flora developed from a disturbance in th ...
in 1963. Sixteen mammalian species including the nine-banded armadillo, the crab-eating raccoon and the red-rumped agouti, 210 species of birds, 24 snakes and 16 lizards have been recorded from the area. The
ichthyofauna Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
is limited - only four of Tobago's 13 freshwater fish species are found in its rivers and streams. The spotted algae-eating goby, '' Sicydium punctatum'', is the most widespread fish species in the area, while the jumping guabine, ''
Anablepsoides hartii ''Anablepsoides'' is a genus of killifish in the family Rivulidae native to tropical South America and the Lesser Antilles. The majority are from the Amazon and Orinoco basins, as well as freshwater systems in the Guiana Shield, but a few species ...
'', is found at the highest elevations and is the only fish species upstream of several waterfalls. Aquatic invertebrates include the decapods ''Atya innocous'', ''Eudaniela garmani'', ''
Macrobrachium faustinum ''Macrobrachium faustinum'' is a species of freshwater shrimp Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod or ...
'', and the introduced
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
'' Melanoides tuberculata.''


Conservation

The Main Ridge Forest Reserve, a
forest reserve A nature reserve (also known as a wildlife refuge, wildlife sanctuary, biosphere reserve or bioreserve, natural or nature preserve, or nature conservation area) is a protected area of importance for flora, fauna, or features of geological or ...
, is one of the oldest protected areas in the world. The Trinidad and Tobago government submitted the Main Ridge Forest Reserve as a tentative listing for a
UNESCO World Heritage Site A World Heritage Site is a landmark or area with legal protection by an international convention administered by the United Nations Educational, Scientific and Cultural Organization (UNESCO). World Heritage Sites are designated by UNESCO for h ...
in 2011. The area is managed by the Department of Natural Resources and the Environment, a department of the Tobago House of Assembly. It has been identified as one of seven
Important Bird Area An Important Bird and Biodiversity Area (IBA) is an area identified using an internationally agreed set of criteria as being globally important for the conservation of bird populations. IBA was developed and sites are identified by BirdLife Int ...
s in Trinidad and Tobago by
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
. The Main Ridge Forest Reserve plays an important role in the protection of native biodiversity, particularly endemic plant species. The Main Ridge and the Northern Range (in northern
Trinidad Trinidad is the larger and more populous of the two major islands of Trinidad and Tobago. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is often referred to as the southernmos ...
) are the areas in the country which support the largest numbers of globally rare plant species. A gap analysis of
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
vascular plants of Trinidad and Tobago identified 15 endemic plant species in the area;
species distribution modelling Species distribution modelling (SDM), also known as environmental (or ecological) niche modelling (ENM), habitat modelling, predictive habitat distribution modelling, and range mapping uses computer algorithms to predict the distribution of a spec ...
suggested that the reserve included suitable habitat for another nine plant endemic species. Main Ridge provides critical habitat for two endemic amphibians, ''
Pristimantis turpinorum ''Pristimantis turpinorum'' is a species of frog in the family Strabomantidae. It is endemic to the north-eastern part of the island of Tobago in The Republic of Trinidad and Tobago. Its natural habitat is tropical moist lowland forest ...
'' (Turpin's frog) and '' Mannophryne olmonae'' (Bloody Bay tree frog), which are endemic to northeastern Tobago and for the coral snake mimic ''
Erythrolamprus pseudoreginae ''Erythrolamprus pseudoreginae'', the Tobago stream snake, is a species of snake in the family Colubridae. The species is found in the northeastern forests of the island of Tobago. Etymology The species name ''pseudoreginae'' was chosen because ...
''. The
white-tailed sabrewing The white-tailed sabrewing (''Campylopterus ensipennis'') is a Near Threatened species of hummingbird in the "emeralds", tribe Trochilini of subfamily Trochilinae. It is found on Tobago and in Venezuela.HBW and BirdLife International (2021) Han ...
(''Campylopterus ensipennis'') is endemic to northeastern Venezuela and the Main Ridge; after Hurricane Flora in 1963 the hummingbird was thought to be extirpated from Tobago, but was rediscovered in 1974. Plant species endemic to Tobago which have been recorded from Main Ridge include '' Odontonema brevipes'', ''
Duguetia tobagensis ''Duguetia tobagensis'' is a small tree in the plant family Annonaceae which is endemic to Trinidad and Tobago. The species is only known from Tobago. Description ''Dugetia tobagensis'' is a small tree, the height of which is unknown. The lea ...
'', '' Phyllanthus acacioides'', '' Besleria seitzii'', '' Cybianthus pittieri'', '' Pilea tobagensis'', and ''
Justicia tobagensis ''Justicia tobagensis'' is a species of plant in the family Acanthaceae which is endemic to Trinidad and Tobago. The species is only known from two areas in the Main Ridge of Tobago. It was first described as ''Drejerella tobagensis'' by Germ ...
''.


Ecotourism

Main Ridge is an important ecotourism destination and birdwatching site. The site has a visitor centre and a network of nine trails which allow access to the site. Additional trails were constructed to reduce pressure on the most popular trail, Gilpin Trace, which is prone to over-use. The Main Ridge Forest Reserve was named the World's Leading Ecotourism Destination by the World Travel Awards from 2003 to 2006 and the World's Leading Green Destination in 2007 and 2009.


Threats

Forests on the Main Ridge were damaged by Hurricane Flora in 1963, and its forests continue to be susceptible to hurricane damage. Given the small size of the area, fires started in surrounding agricultural lands can penetrate deep into the hills. Commercial and subsistence hunting and overuse for tourism pose threats to biodiversity, while the chytrid fungus '' Batrachochytrium dendrobatidis'' poses a threat to endemic amphibians.


References

{{Use dmy dates, date=July 2019 Tobago Mountain ranges of Trinidad and Tobago Protected areas of Trinidad and Tobago