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The yellow-faced grassquit (''Tiaris olivaceus'') is a
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
in the tanager family Thraupidae and is the only member of the genus ''Tiaris''. It is native to the
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
,
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, and the
Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean Se ...
.


Taxonomy

In 1760 the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published works ...
included a description of the yellow-faced grassquit in his ''Ornithologie'' based on a specimen collected in
Saint-Domingue Saint-Domingue () was a French colony in the western portion of the Caribbean island of Hispaniola, in the area of modern-day Haiti, from 1659 to 1804. The name derives from the Spanish main city in the island, Santo Domingo, which came to refer ...
(now
Haiti Haiti (; ht, Ayiti ; French: ), officially the Republic of Haiti (); ) and formerly known as Hayti, is a country located on the island of Hispaniola in the Greater Antilles archipelago of the Caribbean Sea, east of Cuba and Jamaica, and ...
). He used the French name ''Le bruant de S. Domingue'' and the Latin name ''Emberiza dominicensis''. The two stars (**) at the start of the paragraph indicates that Brisson based his description on the examination of a specimen. Although Brisson coined Latin names, these do not conform to the
binomial system The binomial system ( es, Sistema binominal) is a voting system that was used in the legislative elections of Chile between 1989 and 2013. From an electoral system point of view, the binomial system is in effect the D'Hondt method with an ope ...
and are not recognised by the
International Commission on Zoological Nomenclature The International Commission on Zoological Nomenclature (ICZN) is an organization dedicated to "achieving stability and sense in the scientific naming of animals". Founded in 1895, it currently comprises 26 commissioners from 20 countries. Orga ...
. When in 1766 the Swedish naturalist
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
updated his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
'' for the twelfth edition he added 240 species that had been previously described by Brisson. One of these was the yellow-faced grassquit. Linnaeus included a terse description, coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Emberiza olivacea'' and cited Brisson's work. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''olivaceus'' is New Latin for "olive-green". This is now the only species placed in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Tiaris'' that was introduced by the English naturalist
William John Swainson William John Swainson FLS, FRS (8 October 1789 – 6 December 1855), was an English ornithologist, malacologist, conchologist, entomologist and artist. Life Swainson was born in Dover Place, St Mary Newington, London, the eldest son of ...
in 1827. The genus formerly contained additional species. A molecular phylogenetic study published in 2014 found that the genus was non-
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...
and in the subsequent rearrangement four species were moved to other genera. The yellow-faced grassquit was traditionally placed with the buntings and New World sparrows in the family Emberizidae, but
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
studies have shown that it is a member of the tanager family Thraupidae and belongs to the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Coerebinae that also contains
Darwin's finches Darwin's finches (also known as the Galápagos finches) are a group of about 18 species of passerine birds. They are well known for their remarkable diversity in beak form and function. They are often classified as the subfamily Geospizinae or t ...
. Five
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised: * ''T. o. pusillus'' Swainson, 1827 – east Mexico to Ecuador and Venezuela * ''T. o. intermedius'' ( Ridgway, 1885) – island of Cozumel (off southeast Mexico) * ''T. o. ravidus'' Wetmore, 1957 –
Coiba Island Coiba is the largest island in Central America, with an area of , off the Pacific coast of the Panamanian province of Chiriquí. It is part of the Tolé District of that province. History Coiba separated from continental Panama about 12,000 to ...
(off south Panama) * ''T. o. olivaceus'' (Linnaeus, 1766) – Cuba,
Isla de la Juventud Isla de la Juventud (; en, Isle of Youth) is the second-largest Cuban island (after Cuba's mainland) and the seventh-largest island in the West Indies (after mainland Cuba itself, Hispaniola, Jamaica, Puerto Rico, Trinidad, and Andros Islan ...
,
Cayman Islands The Cayman Islands () is a self-governing British Overseas Territory—the largest by population in the western Caribbean Sea. The territory comprises the three islands of Grand Cayman, Cayman Brac and Little Cayman, which are located to the ...
, Jamaica and Hispaniola * ''T. o. bryanti'' (Ridgway, 1898) – Puerto Rico and nearby islands


Description

It is a small bird with a conical bill, sharper than that of the related
seedeater {{Unreferenced, date=December 2007 The seedeaters are a form taxon of seed-eating passerine birds with a distinctively conical bill. Most are Central and South American birds that were formerly placed in the American sparrow family (Passerell ...
s. It is long and weighs about , depending on
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
. The adult male has an olive-green back, and its face and breast are black apart from a bright yellow throat,
supercilia The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also ...
, and lower eyelid spot. The rest of the underparts are greyish olive. The
beak The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food ...
and eyes are dark, the legs are grey. The adult female is slightly smaller on average than the male. It is dull olive-green above and paler grey below, and may have some dark breast smudges. The yellow face pattern is much weaker and duller, and may be almost invisible. The lower part of the beak is dark horn-colored (light grey). Young birds are coloured essentially like the adult female, but duller and greyer. Young males begin to acquire full adult plumage in their first year. The yellow-faced grassquit has a weak buzzing trilled ''ttttt-tee'' call. The song is a varying series of high thin rapid trills. Given for a prolonged time, it is melodious, yet subdued, and often only heard from a short distance away.


Distribution and ecology

It breeds from central
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
to northern
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...
and north-western
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
, and also on the
Greater Antilles The Greater Antilles ( es, Grandes Antillas or Antillas Mayores; french: Grandes Antilles; ht, Gwo Zantiy; jam, Grieta hAntiliiz) is a grouping of the larger islands in the Caribbean Sea, including Cuba, Hispaniola, Puerto Rico, Jamaica, and ...
and nearby islands. It is not a migratory bird but moves about outside the breeding season; vagrants have been recorded in Southern Texas and Florida. It has been introduced to
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
. The yellow-faced grassquit is a common to abundant resident in lowlands and foothills up to altitude in semi-open areas such as roadsides, pasture, weedy fields and low scrub. It avoids dry grassland and other very low growth, but will readily utilize neglected gardens where lawn grass has grown high. It is a fairly sociable bird, sometimes forming loose flocks with other tanagers (such as
seedeater {{Unreferenced, date=December 2007 The seedeaters are a form taxon of seed-eating passerine birds with a distinctively conical bill. Most are Central and South American birds that were formerly placed in the American sparrow family (Passerell ...
s) and emberizids that share its lifestyle. Occasionally, adult males will come together to perch in a shrub and sing. This species feeds mainly on grass seeds, but also takes other seeds, berries and some insects. Food is typically plucked from the inflorescences directly, or gathered from the ground. Animal prey is hunted among foliage in a New World warbler-like fashion. The main breeding season is in the summer months, but except for February to April birds may nest at other times of the year. This species sometimes forms loose nesting colonies. During courtship, the male vibrates his wings as he sings his subdued song, sitting only away so the female can properly hear him. The roughly globular nest, built by the female, is made of grass and weed stems compacted into a thick mass, and lined with pieces of grass inflorescences and bast fibre. It has a side entrance and is placed usually less than above the ground, often among grass or weeds on a road or river
embankment Embankment may refer to: Geology and geography * A levee, an artificial bank raised above the immediately surrounding land to redirect or prevent flooding by a river, lake or sea * Embankment (earthworks), a raised bank to carry a road, railwa ...
. Occasionally, the species nests in shrubs up to above the ground. The
clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts). ...
is two or three, rarely four, brown-speckled white eggs. They are incubated by the female alone for 12–14 days to hatching. This bird is not rare and widely distributed; it is thus classified a species of least concern by the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
. It seems to benefit from deforestation, increasing in numbers and expanding its range; for example, it is only since 1997 known from the eastern Andean slope of Meta Department in
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
.


References


Further reading

*


External links

* * * * {{Taxonbar, from=Q728336 Birds of the Greater Antilles Birds of Mexico Birds of Central America Birds of Colombia
yellow-faced grassquit The yellow-faced grassquit (''Tiaris olivaceus'') is a passerine bird in the tanager family Thraupidae and is the only member of the genus ''Tiaris''. It is native to the Central America, South America, and the Caribbean. Taxonomy In 1760 the ...
Taxa named by Carl Linnaeus