Tiaris Bicolor
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The black-faced grassquit (''Melanospiza bicolor'') is a small
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
. It is recognized as a
tanager The tanagers (singular ) comprise the bird family Thraupidae, in the order Passeriformes. The family has a Neotropical distribution and is the second-largest family of birds. It represents about 4% of all avian species and 12% of the Neotropica ...
closely related to
Darwin's finches Darwin's finches (also known as the Galápagos finches) are a group of about 18 species of passerine birds. They are well known for their remarkable diversity in beak form and function. They are often classified as the subfamily Geospizinae or t ...
. It breeds in the
West Indies The West Indies is a subregion of North America, surrounded by the North Atlantic Ocean and the Caribbean Sea that includes 13 independent island countries and 18 dependencies and other territories in three major archipelagos: the Greater A ...
except
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribbea ...
, on
Tobago Tobago () is an List of islands of Trinidad and Tobago, island and Regions and municipalities of Trinidad and Tobago, ward within the Trinidad and Tobago, Republic of Trinidad and Tobago. It is located northeast of the larger island of Trini ...
but not
Trinidad Trinidad is the larger and more populous of the two major islands of Trinidad and Tobago. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is often referred to as the southernmos ...
, and along the northern coasts of
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
and
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
.


Taxonomy

The first formal description of the black-faced grassquit was by the Swedish naturalist
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in 1766 in the twelfth edition of his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
''. He introduced the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Fringilla bicolor''. Linnaeus based his description on the "Bahama Sparrow" that was described and illustrated by
Mark Catesby Mark Catesby (24 March 1683 – 23 December 1749) was an English naturalist who studied the flora and fauna of the New World. Between 1729 and 1747 Catesby published his ''Natural History of Carolina, Florida and the Bahama Islands'', the fi ...
in his ''The Natural History of Carolina, Florida and the Bahama Islands'' which was published between 1729 and 1732. The black-faced grassquit was traditionally placed in the genus ''
Tiaris The yellow-faced grassquit (''Tiaris olivaceus'') is a passerine bird in the tanager family Thraupidae and is the only member of the genus ''Tiaris''. It is native to the Central America, South America, and the Caribbean. Taxonomy In 1760 the F ...
''. A
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
study published in 2014 found that this genus was
polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of converg ...
and that the black-faced grassquit was closely related to the
Saint Lucia black finch The Saint Lucia black finch (''Melanospiza richardsoni'') is a species of bird in the family Thraupidae. It is endemic to Saint Lucia, where its natural habitats are subtropical or tropical moist lowland forest, subtropical or tropical dry shru ...
in the
monospecific In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
genus ''
Melanospiza ''Melanospiza'' is a genus of Neotropical birds in the tanager family Thraupidae. Taxonomy and species list The genus ''Melanospiza '' was introduced in 1897 by the American ornithologist Robert Ridgway with the Saint Lucia black finch as the ty ...
''. In the resulting reorganization of the genera, the black-faced grassquit was moved to ''Melanospiza''. Eight
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised: * ''M. b. bicolor'' (Linnaeus, 1766) – Bahamas and islands off north Cuba * ''M. b. marchii'' ( Baird, SF, 1864) – Jamaica and Hispaniola * ''M. b. omissa'' (
Jardine Jardine is a surname. Notable people with the surname include: * Al Jardine (born 1942), member of the Beach Boys * Alexander Jardine (Medal of Honor) (1874–1949), American Medal of Honor recipient * Antonio Jardine (born 1988), NCAA college bas ...
, 1847) – Puerto Rico, Lesser Antilles, north Colombia and north Venezuela * ''M. b. huilae'' ( Miller, AH, 1952) – central Colombia * ''M. b. grandior'' (
Cory As a given name, Cory is used by both males and females. It is a variation of the name Cora, which has Greek origins and is the maiden name of the goddess Persephone. The name also can have origins from the Gaelic word ''coire'', which means "in ...
, 1887) – San Andrés Archipelago (east of Nicaragua) * ''M. b. johnstonei'' ( Lowe, 1906) –
La Blanquilla Island Blanquilla is an island, one of the Federal Dependencies of Venezuela, Federal Dependencies of Venezuela, located in the southeastern Caribbean Sea about 293 km (182 miles) northeast of Caracas. It is a popular location for divers, as well ...
and the
Los Hermanos Archipelago The Los Hermanos Archipelago is a chain of seven rocky barren islets that is part of the Federal Dependencies of Venezuela, at .Hernández Caballero, Serafín (Editor). 1998: Gran Enciclopedia de Venezuela. Editorial Globe, C.A. Caracas. 10 volúme ...
(off north Venezuela) * ''M. b. sharpei'' (
Hartert Ernst Johann Otto Hartert (29 October 1859 – 11 November 1933) was a widely published German ornithologist. Life and career Hartert was born in Hamburg, Germany on 29 October 1859. In July 1891, he married the illustrator Claudia Bernadine E ...
, 1893) – Netherlands Antilles * ''M. b. tortugensis'' (Cory, 1909) –
La Tortuga Island La Tortuga Island (; "La Tortuga" means "the turtle") is an uninhabited island of Venezuela, the largest in the Federal Dependencies of Venezuela. It is part of a group of islands that include the Tortuguillos and Cayo Herradura. Isla La Tortu ...
(off north Venezuela)


Description

A male black-faced grassquits is around long and weighs approximately . It has a short conical black bill, a black head and breast with an olive green back. Females and immature birds have dull olive-grey upperparts and head, and paler grey underparts becoming whiter on the belly. Males on the
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
n mainland have more extensively black underparts, shading to a grey belly. The male has a display flight in which he flies for short distances, vibrating his wings and giving a buzzing ''dik-zeezeezee'' call.


Behaviour and ecology


Breeding

This is a common bird in long grass or scrub in open or semi-open areas, including roadsides and
rice Rice is the seed of the grass species ''Oryza sativa'' (Asian rice) or less commonly ''Oryza glaberrima ''Oryza glaberrima'', commonly known as African rice, is one of the two domesticated rice species. It was first domesticated and grown i ...
fields. It makes a domed grass nest, lined with finer grasses, and placed low in a bush or on a bank. The typical clutch is two or three whitish eggs blotched with reddish brown.


Food and feeding

The black-faced grassquit feeds mainly on seeds, especially of grasses and weeds, occasionally on fruits and berries.https://sta.uwi.edu/fst/lifesciences/sites/default/files/lifesciences/documents/ogatt/Tiaris_bicolor%20-%20Black-faced%20Grassquit.pdf It also feeds on small insects, mainly during the breeding season. It is often found in small groups.


References


External links


Xeno-canto: audio recordings of the black-faced grassquit

Black-faced Grassquit videos, photos & sounds
on the Internet Bird Collection {{Taxonbar, from=Q941987 black-faced grassquit Birds of the Caribbean Birds of the Dominican Republic Birds of Colombia Birds of Venezuela black-faced grassquit black-faced grassquit