The Ringlet
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The ringlet (''Aphantopus hyperantus'') is a butterfly in the family Nymphalidae. It is only one of the numerous "ringlet" butterflies in the tribe Satyrini.


Range

The ringlet is a widely distributed species found throughout much of the Palearctic realm. In Europe it is common in most countries but absent from northern Scandinavia, peninsular Italy (found in northern Italy), Portugal, southern and central Spain (found in Cantabrian Mountains and the eastern Pyrenees), the Mediterranean islands and North Africa. In Greece it is found in northern regions (
Macedonia Macedonia most commonly refers to: * North Macedonia, a country in southeastern Europe, known until 2019 as the Republic of Macedonia * Macedonia (ancient kingdom), a kingdom in Greek antiquity * Macedonia (Greece), a traditional geographic reg ...
, Thessaly). Beyond Europe it is found across much of temperate Asia including Russia, Siberia, Mongolia,
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
and Korea.


Description

''Aphantopus hyperantus'' is a medium-sized butterfly with a wingspan of up to 35 to 42 millimeters. The wing upper and lower sides are solid brown with small, yellowish-rimmed eyespots. The newly emerged ringlet has a velvety appearance and is almost black with a white fringe to the wings. The number and size of the eyespots is variable, they may be missing on the upper wing surface. In central Europe and southern England the rare form ''arete'' occurs. The eggs are pale yellow when first laid, but become pale brown. The caterpillars are about 25 millimeters long. They are gray or light reddish brown and have dark, reddish brown and very fine dots. Dorsally there is a dark longitudinal line, which is widened at the segment boundaries. Toward the rear, this line is more intensely colored. The head is darker and has several faint longitudinal stripes.


Color and wing spot variation

Ringlet butterfly (Aphantopus hyperantus) 1 spot newly emerged.jpg, Newly emerged, with one spot Ringlet butterfly (Aphantopus hyperantus) 1 spot.jpg, Fresh, with one spot Ringlet butterfly (Aphantopus hyperantus) 2 spots worn.jpg, Worn specimen with two spots Ringlet butterfly (Aphantopus hyperantus) 3 spots.jpg, Faded specimen with three spots


Subspecies

*ssp. ''abaensis'' Yoshino, 2003 - northwestern Sichuan *ssp. ''alpheois'' Fruhstorfer, 1908 - Ural, western Siberia *ssp. ''arctica'' (Seitz, 1909) - northern Europe *ssp. ''bieti'' (Oberthür, 1884) - Sichuan and northern Yunnan *ssp. ''hyperantus'' Linnaeus, 1758 - western Europe, the type locality is Sweden *ssp. ''luti'' Evans, 1915 - southeastern Tibet *ssp. ''ocellata'' (Butler, 1882) (= ''amurensis'' Staudinger, 1892; = ''insularis'' Kurentzov, 1966) - Amur and Ussuri *ssp. ''sajana'' (O. Bang-Haas, 1906) - Sayan Mountains *ssp. ''sibiricus'' Obraztsov, 1936 - Altai, southern Siberia and Transbaikalia


Habitat

They live in grassy, moist or dry forest clearings with bushes but not in open places. There is a strong degree of attachment to woodland edges and blackberry bushes. The insect can also be very common where there are creeping thistles ('' Cirsium arvense'') or swamp thistles ('' Cirsium palustre''), oregano ('' Origanum vulgare''), forest scabious (''
Knautia sylvatica ''Knautia'' is a genus of flowering plants in the family Caprifoliaceae. The common names are variants of "widow flower". Others are given the name "scabious", which properly belongs to the related genus ('' Scabiosa''). The name ''Knautia'' com ...
''), or hogweed ('' Heracleum sphondylium'') which are favorite food plants of the imagos. The males fly in search of newly hatched females in slow, uninterrupted flight and flutter round, about and between grass stems.


Flight period

A single brood butterfly, the imagines fly from mid-June to late August.


Food of the larva

The caterpillars feed on many grasses. Among the food plants are:


Development and biology

The female scatters non-adhesive eggs in a slow low flight over grasslands. The larva is nocturnal. There are four moults. The larva hibernates while in the third instar, breaking diapause to feed on warm winter evenings. Feeding resumes in the spring. The pupa stands generally upright in a flimsy silk cocoon, at the base of a grass tussock. This stage lasts for two weeks. ''A. hyperantus'' is generally considered to have a closed population structure since it occurs in small, well-defined populations.


Etymology

Hyperantus, of Greek mythology, was one of the 50 sons of
Aegyptus In Greek mythology, Aegyptus or Ægyptus (; grc, Αἴγυπτος) was a legendary king of ancient Egypt. He was a descendant of the princess Io through his father Belus, and of the river-god Nilus as both the father of Achiroe, his mother ...
, killed by one of the 50 daughters of Danaus.


References

*Tom Tolman, Richard Lewington ''The Butterflies of Europe and Northwest Africa''. Nabu-Kosmos, Stuttgart 1998, *Heiko Bellmann ''Der neue Kosmos-Schmetterlingsführer, Schmetterlinge, Raupen und Futterpflanzen''. Franckh-Kosmos, Stuttgart 2003ISBN 3-440-09330-1 *Günter Ebert, Erwin Rennwald (Hrsg.) Tagfalter. 2. Spezieller Teil: Satyridae, Libytheidae, Lycaenidae, Hesperiidae. In: ''Die Schmetterlinge Baden-Württembergs''. 1. Auflage. Band 2, Ulmer, Stuttgart (Hohenheim) 1991, .


External links


Eurobutterflies Matt Rowlings
* ttp://www.ukbutterflies.co.uk/species.php?species=hyperantus UK Butterflies {{Taxonbar, from=Q161520 Aphantopus Butterflies of Asia Butterflies of Europe Butterflies described in 1758 Taxa named by Carl Linnaeus Taxobox binomials not recognized by IUCN