Teucrium Cossonii Gamble
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''Teucrium'' is a
cosmopolitan Cosmopolitan may refer to: Food and drink * Cosmopolitan (cocktail), also known as a "Cosmo" History * Rootless cosmopolitan, a Soviet derogatory epithet during Joseph Stalin's anti-Semitic campaign of 1949–1953 Hotels and resorts * Cosmopoli ...
genus of flowering plants in the family
Lamiaceae The Lamiaceae ( ) or Labiatae are a family of flowering plants commonly known as the mint, deadnettle or sage family. Many of the plants are aromatic in all parts and include widely used culinary herbs like basil, mint, rosemary, sage, savory ...
, commonly known as germanders. Plants in this genus are
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
herbs In general use, herbs are a widely distributed and widespread group of plants, excluding vegetables and other plants consumed for macronutrients, with savory or aromatic properties that are used for flavoring and garnishing food, for medicinal ...
or shrubs, with branches that are more or less square in cross-section, leaves arranged in opposite pairs, and flowers arranged in
thyrse A thyrse is a type of inflorescence in which the main axis grows indeterminately, and the subaxes (branches) have determinate growth.{{cite book , author1=Hickey, M., author2=King, C. , year=2001 , title=The Cambridge Illustrated Glossary of Bot ...
s, the corolla with mostly white to cream-coloured, lobed petals.


Description

Plants in the genus ''Teucrium'' are perennial herbs or shrubs with four-cornered stems, often with simple hairs and
sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...
glands In animals, a gland is a group of cells in an animal's body that synthesizes substances (such as hormones) for release into the bloodstream (endocrine gland) or into cavities inside the body or its outer surface (exocrine gland). Structure De ...
. The leaves are arranged in opposite pairs, simple or with three leaflets sometimes with lobed or serrated edges. The flowers are arranged in a thyrse, sometimes in a cyme in leaf axils. The flowers have five more or less similar
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
s fused at the base, and the corolla is white or cream-coloured with five lobes forming two lips. The upper lip is usually much reduced in size and the lower lip has three lobes, the central lobe usually larger than the side lobes. There are four
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s attached near the base of the petals and the fruit is a
schizocarp A schizocarp is a dry fruit that, when mature, splits up into mericarps. There are different definitions: * Any dry fruit composed of multiple carpels that separate. : Under this definition the mericarps can contain one or more seeds (the m ...
with four segments.


Taxonomy

The genus ''Teucrium'' was first formally described in 1753 by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in ''
Species Plantarum ' (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genera. It is the first work to consistently apply binomial names and was the ...
''. The name ''Teucrium'' was used by
Pedanius Dioscorides Pedanius Dioscorides ( grc-gre, Πεδάνιος Διοσκουρίδης, ; 40–90 AD), “the father of pharmacognosy”, was a Greek physician, pharmacologist, botanist, and author of ''De materia medica'' (, On Medical Material) —a 5-vol ...
for several species in this genus, and is believed to refer to
King Teucer In Greek mythology, King Teucer (; Ancient Greek: Τεῦκρος ''Teûkros'') was said to have been the son of the river-god Scamander and the nymph Idaea. Mythology Before the arrival of Dardanus, the land that would eventually be called D ...
of
Troy Troy ( el, Τροία and Latin: Troia, Hittite language, Hittite: 𒋫𒊒𒄿𒊭 ''Truwiša'') or Ilion ( el, Ίλιον and Latin: Ilium, Hittite language, Hittite: 𒃾𒇻𒊭 ''Wiluša'') was an ancient city located at Hisarlik in prese ...
who used the plant in his medicine.


Species

(See also List of ''Teucrium'' species) ''Teucrium'' is a cosmopolitan genus with about 300 species, the distribution centred on the Mediterranean. There are about thirteen species endemic to Australia.


Selected species

* ''
Teucrium albicaule ''Teucrium albicaule'', commonly known as scurfy germander, is a species of flowering plant in the family Lamiaceae, and is endemic to inland areas of Australia. It is a hoary, perennial herb that spreads by root suckers, forming dense stands. ...
''
Toelken Toelken is a surname. Notable people with this surname include: *Barre Toelken (1935–2018), American folklorist *Ernst Heinrich Toelken (1795–1878), German anatomist, physicist, and archaeologist *Hellmut R. Toelken Hellmut R. Toelken (born 19 ...
– scurfy germander (Australia) * ''
Teucrium arduinoi ''Teucrium'' is a Cosmopolitan distribution, cosmopolitan genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are perennial plant, perennial herbaceous plant, herbs or shrubs, with branches that a ...
'' L. * ''
Teucrium argutum ''Teucrium argutum'', commonly known as native germander, is a species of flowering plant in the family Lamiaceae, and is endemic to eastern Australia. It is a perennial herb often suckering, with hairy, broadly egg-shaped leaves with toothed or ...
'' R.Br. – native germander (Qld., N.S.W.) * ''
Teucrium aroanium ''Teucrium aroanium'' is a species of flowering plant in the family Lamiaceae. It is endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined z ...
'' Orph. ex Boiss. * ''
Teucrium balfourii ''Teucrium balfourii'' is a species of flowering plant in the family Lamiaceae. It is found only in the Socotra Islands, part of the nation of Yemen. Its natural habitat In ecology, the term habitat summarises the array of resources, phy ...
'' Vierh. * ''
Teucrium balthazaris ''Teucrium balthazaris'' is a species of flowering plant in the family Lamiaceae. It is found only in Spain. Its natural habitat is Mediterranean-type shrubby vegetation. It is threatened by habitat loss Habitat destruction (also termed hab ...
'' Sennen * ''
Teucrium betonicum ''Teucrium'' is a cosmopolitan genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are perennial herbs or shrubs, with branches that are more or less square in cross-section, leaves arranged in op ...
'' L'Hér. * ''
Teucrium botrys ''Teucrium botrys'', the cutleaf germander or cut-leaved germander is a low to short downy annual, sometimes biennial, plant. It was noted by Carl Linnaeus in 1753 and is classified as part of the genus ''Teucrium'' in the family Lamiaceae. It ha ...
'' L. – cut-leaved germander * '' Teucrium canadense'' L. – American germander, Canada germander, wood sage, hairy germander * ''
Teucrium capitatum ''Teucrium'' is a cosmopolitan genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are perennial herbs or shrubs, with branches that are more or less square in cross-section, leaves arranged in op ...
'' L. * ''
Teucrium carolipaui ''Teucrium'' is a Cosmopolitan distribution, cosmopolitan genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are perennial plant, perennial herbaceous plant, herbs or shrubs, with branches that a ...
'' C. Vicioso ex Pau * ''
Teucrium chamaedrys ''Teucrium chamaedrys'', the wall germander, is a species of plant native to the Mediterranean regions of Europe and North Africa, and the Middle East as far as Iran.Dobignard, A. & Chatelain, C. (2012). Index synonymique de la flore d'Afrique d ...
'' L. – wall germander * ''
Teucrium chardonianum ''Teucrium'' is a cosmopolitan genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are perennial herbs or shrubs, with branches that are more or less square in cross-section, leaves arranged in op ...
'' Maire & Wilczek * ''
Teucrium coahuilanum ''Teucrium'' is a cosmopolitan genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are perennial herbs or shrubs, with branches that are more or less square in cross-section, leaves arranged in op ...
'' B.L.Turner * ''
Teucrium corymbosum ''Teucrium corymbosum'', commonly known as forest germander, is a species of flowering plant in the family Lamiaceae and is native to Australia and New Guinea. It is a perennial herb or subshrub densely covered with glands and with narrow egg-sh ...
'' R.Br. – forest germander (Australia, New Guinea) * ''
Teucrium cossonii ''Teucrium cossonii'', the fruity germander, is a perennial herb in the family Lamiaceae, endemic to the Balearic Islands The Balearic Islands ( es, Islas Baleares ; or ca, Illes Balears ) are an archipelago in the Balearic Sea, near t ...
'' D.Wood – fruity teucrium * '' Teucrium creticum'' L. * '' Teucrium cubense'' Jacq. – small coastal germander, dwarf germander * ''
Teucrium eremaeum ''Teucrium eremaeum'' is a species of flowering plant in the family Lamiaceae and is endemic to the south-west of Western Australia. It is a perennial herb or shrub with small, linear to lance-shaped leaves and white or cream-coloured flowers. ...
''
Diels Diels is the last name of several people: * Rudolf Diels (1900–1957), German politician * Otto Diels (1876–1954), German scientist noted for his work on the Diels–Alder reaction * Ludwig Diels (1874–1945), German botanist * Hermann Diels ...
(W.A.) * '' Teucrium divaricatum'' Sieber ex Heldr. * ''
Teucrium flavum ''Teucrium flavum'' is a species of shrub in the family Lamiaceae The Lamiaceae ( ) or Labiatae are a family of flowering plants commonly known as the mint, deadnettle or sage family. Many of the plants are aromatic in all parts and include ...
'' L. * ''
Teucrium fruticans ''Teucrium fruticans'' (common name tree germander or shrubby germander) is a species of flowering plant in the mint family Lamiaceae, native to the western and central Mediterranean. Growing to tall by wide, it is a spreading evergreen shrub w ...
'' L. – tree germander, shrubby germander * ''
Teucrium glandulosum ''Teucrium glandulosum'' is a species of flowering plant in the mint family known by the common names sticky germander and desert germander. It is native to the Sonoran Desert of Arizona, Baja California, Baja California Sur, and San Bernardino ...
'' Kellogg – common germander, desert germander, sticky germander * ''
Teucrium gnaphalodes ''Teucrium gnaphalodes'' is a plant species in the genus ''Teucrium''. It is endemic to the Iberian Peninsula and grows at altitudes between 200 and 1500 m. It flowers from March to July. The flavones diosmin, cirsimaritin, salvigenin, cirsilineo ...
'' L’Hér. * ''
Teucrium grandiusculum ''Teucrium grandiusculum'' is a species of flowering plant in the family Lamiaceae and is endemic to central Australia. It is a perennial herb or shrub with toothed, egg-shaped leaves and white flowers. Description ''Teucrium grandiusculum'' is ...
'' F.Muell. & Tate (W.A., S.A., N.T.) * '' Teucrium heterophyllum'' L’Hér. * ''
Teucrium integrifolium ''Teucrium integrifolium'', commonly known as teucry weed or green germander, is a species of flowering plant in the family Lamiaceae and is endemic to northern Australia. It is a perennial herb with broadly elliptic to broadly egg-shaped leaves ...
'' Benth. – teucry weed (W.A., N.T., Qld.) *''
Teucrium junceum ''Teucrium junceum'' is a species of flowering plant in the family Lamiaceae, and is endemic to eastern Australia. It is a scrambling, openly-branched shrub, with small leaves, white flowers and orange to red fruit. Description ''Teucrium junce ...
'' * ''
Teucrium laciniatum ''Teucrium'' is a cosmopolitan genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are perennial herbs or shrubs, with branches that are more or less square in cross-section, leaves arranged in op ...
'' Torr. – germander, lacy germander * '' Teucrium lepicephalum'' Pau * ''
Teucrium marum ''Teucrium marum'', commonly known as cat thyme or kitty crack, is a species of germander. Its small, oval leaves give it a thyme-like appearance, but the musty scent is quite unlike the delicate aroma of thyme. Cat thyme is a mounding, tender pe ...
'' L. * ''
Teucrium massiliense ''Teucrium'' is a cosmopolitan genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are perennial herbs or shrubs, with branches that are more or less square in cross-section, leaves arranged in op ...
'' L. * ''
Teucrium montanum ''Teucrium montanum'', the mountain germander, is a germander native to southern Europe, from Spain eastward to Turkey. It forms shrubs about a foot high, with small leaves no more than an inch across, and petals blooming yellowish white. On sout ...
'' L. * ''
Teucrium oliverianum ''Teucrium oliverianum'' is a species of flowering plant in the family Lamiaceae, native to Iraq, Kuwait, Saudi Arabia and Iran. It may also possibly occur in Lebanon and Syria. It was first described by George Bentham George Bentham (22 S ...
'' Ging. ex Benth. * ''
Teucrium orientale ''Teucrium'' is a cosmopolitan genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are perennial herbs or shrubs, with branches that are more or less square in cross-section, leaves arranged in op ...
'' L. – oriental germander * ''
Teucrium polium ''Teucrium polium'', known popularly as felty germander, is a sub-shrub and herb native to the western Mediterranean region (Albania, North Macedonia, Spain, France, Algeria, Morocco, Tunisia). Its flowers are small and range from pink to white, ...
'' L. * '' Teucrium pseudochamaepitys'' L. * '' Teucrium puberulum'' * '' Teucrium pyrenaicum'' L. * ''
Teucrium racemosum ''Teucrium racemosum'', also commonly referred to as either the grey germander or forest germander, is a species of flowering plant in the family Lamiaceae. It is endemic to Australia and is found in all mainland states, the Northern Territory ...
'' – forest germander * ''
Teucrium scordium ''Teucrium scordium'' is a species of flowering plant belonging to the family Lamiaceae. Its native range is Europe to China. References

{{Taxonbar, from=Q158832 Teucrium, scordium Flora of Malta ...
'' L. * ''
Teucrium scorodonia ''Teucrium scorodonia'', common name the woodland germander or wood sage, is a species of flowering plant in the genus ''Teucrium'' of the family Lamiaceae. It is native to Western Europe and Tunisia, but cultivated in many places as an ornamenta ...
'' L. – woodland germander * '' Teucrium socotranum'' Vierh. * ''
Teucrium subspinosum ''Teucrium'' is a cosmopolitan genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are perennial herbs or shrubs, with branches that are more or less square in cross-section, leaves arranged in op ...
'' Pourr. ex Willd. * '' Teucrium teucriiflorum'' (F.Muell.) Kattari & Salmaki * '' Teucrium townsendii'' ** ''Teucrium townsendii'' ssp. ''affine'' ** ''Teucrium townsendii'' var. ''townsendii'' * '' Teucrium vesicarium'' Mill. * ''
Teucrium viscidum ''Teucrium'' is a cosmopolitan genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are perennial herbs or shrubs, with branches that are more or less square in cross-section, leaves arranged in op ...
'' Blume * ''
Teucrium werneri ''Teucrium'' is a cosmopolitan genus of flowering plants in the family Lamiaceae, commonly known as germanders. Plants in this genus are perennial herbs or shrubs, with branches that are more or less square in cross-section, leaves arranged in op ...
'' Emb.


Fossil record

†''Teucrium tatjanae'' seed
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s are known from the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
,
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
and
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
, Miocene and Pliocene of central and southern
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
and Miocene of
Lusatia Lusatia (german: Lausitz, pl, Łużyce, hsb, Łužica, dsb, Łužyca, cs, Lužice, la, Lusatia, rarely also referred to as Sorbia) is a historical region in Central Europe, split between Germany and Poland. Lusatia stretches from the Bóbr ...
. The fossil seeds are similar to seeds of the extant ''Teucrium orientale''. †''Teucrium pripiatense'' seed fossils have been described from the Pliocene Borsoni Formation in the
Rhön Mountains The Rhön Mountains () are a group of low mountains (or ''Mittelgebirge'') in central Germany, located around the border area where the states of Hesse, Bavaria and Thuringia come together. These mountains, which are at the extreme southeast end o ...
of central
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
.The floral change in the tertiary of the Rhön mountains (Germany) by Dieter Hans Mai - Acta Paleobotanica 47(1): 135-143, 2007.


References


External links


USDA PLANTS Profile
{{Taxonbar, from=Q157714 * Lamiaceae genera Taxa named by Carl Linnaeus