Tetratheca Pilosa
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''Tetratheca pilosa'' is a flowering plant in the family
Elaeocarpaceae Elaeaocarpaceae is a family of flowering plants. The family contains approximately 615 species of trees and shrubs in 12 genera."Elaeocarpaceae" In: Klaus Kubitzki (ed.). ''The Families and Genera of Vascular Plants'' vol. VI. Springer-Verlag: Be ...
, endemic to Australia. It is a small shrub found in dry sclerophyll forests, open heathlands and woodlands of Australia. It was first recorded in 1805 by French botanist
Jacques Labillardière Jacques-Julien Houtou de Labillardière (28 October 1755 – 8 January 1834) was a French biologist noted for his descriptions of the flora of Australia. Labillardière was a member of a voyage in search of the Jean-François de Galaup, comte ...
.


Description

Characterized by pink to mauve drooping (pendulous), auxiliary, solitary flowers, which are radially symmetrical, consisting of 4 sepals and 4 petals, 6 mm long. Stamens usually 8, opening by an apical pore and form a dark center to the flower above a superior ovary. Stamen tube widest between the base and the apex and are often hidden by the petals, hence the common name black eyed susan. Stems are erect, unbranched or branched from the base. Leaves are green, narrow, distinctly alternate and slightly revolute or with recurved margins. Leaves can be hairless or have non-glandular hairs. Up to 15 mm in length. ''Tetratheca'' fruit have locules that dehisce as the fruit desiccates, releasing 1 to 5 seeds.


Distribution

''Tetratheca pilosa'' is one of a few ''Tetratheca'' species that occurs in multiple states across Australia (most other species are locally endemic). It is found in Tasmania, Victoria, the south east corner of South Australia and in a few localised patches in New South Wales.


Ecology

The species exists in open woodland, heathland and dry sclerophyll forests. The vegetation structure generally consists of a sparse understorey and an open canopy of ''Eucalyptus''. Depending on substrate the canopy trees may be dominated by ''E. amygdalina'', ''E. delegatensis'' or ''E. obliqua.'' A sparse understorey of ''Banksia marginata'' and ''Exocarpus cupressiformis'' is common. ''T. pilosa'' grows sparsely amongst other woody shrub species such as ''Epacris impressa'', ''Pultenaea juniperina'', ''Davisea latifolia.'' Ground cover usually consists of a sparse layer of ''Gonocarpus teucroides'' and a high coverage of fine litter and rocks. ''T. pilosa'' does poorly when competing with other plants, but has a strong advantage in sandy, gravely, hydrophobic, acidic and nutrient poor soils. It persists on sandstone,
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...
mudstone and siltstone and soils of granitic origins. The black centre creates a target for native bees which are capable of
buzz pollination Buzz pollination or sonication is a technique used by some bees, such as solitary bees to release pollen which is more or less firmly held by the anthers. The anthers of buzz-pollinated plant species are typically tubular, with an opening at onl ...
. Buzz pollination requires the bee to use their flight mascles to vibrate the pollen loose, a skill which European bees do not have. Only 9% of the world plants use this form of pollination.  ''Tetratheca'' seeds are a food source for ants which collect, disperse and take the seed underground. This has multiple advantages for the plant, creating a soil seed store, protected from fire and dispersed further than wind may be able to carry the seed.


References

{{Taxonbar, from=Q18009907
pilosa The order Pilosa is a clade of xenarthran placental mammals, native to the Americas. It includes the anteaters and sloths (which includes the extinct ground sloths). The name comes from the Latin word for "hairy". Origins and taxonomy The b ...
Taxa named by Jacques Labillardière Plants described in 1805