''Tetraphalerus'' is a genus of

beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...

s in the family

Ommatidae The Ommatidae are a family of beetles in the suborder Archostemata. The Ommatidae are considered the extant beetle family that has most ancestral characteristics. There are only seven extant species, confined to Australia and South America. How ...

, It is currently known from two extant species native to South America and several fossil species from the Jurassic and Cretaceous of Asia.

Systematics

According to Kirejtshuk, 2020. * †'' Tetraphalerus brevis'' Ponomarenko, 1964

Karabastau Formation The Karabastau Formation ( kk, Qarabastaý svıtasy) is a geological formation and lagerstätte in the Karatau Mountains of southern Kazakhstan whose strata date to the Middle to Late Jurassic. It is an important locality for insect fossils that ...

, Kazakhstan, Oxfordian *'' Tetraphalerus bruchi'' Heller, 1913, recent, Northern Argentina * †'' Tetraphalerus collaris'' Ponomarenko, 1997 Dzun-Bain Formation, Mongolia,

Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...

* †'' Tetraphalerus glabratus'' Ponomarenko, 1997 Togo-Khuduk Formation, Mongolia,

Bajocian In the geologic timescale, the Bajocian is an age and stage in the Middle Jurassic. It lasted from approximately 170.3 Ma to around 168.3 Ma (million years ago). The Bajocian Age succeeds the Aalenian Age and precedes the Bathonian Age. Stratig ...

Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Strat ...

* †'' Tetraphalerus largicoxa'' Lin Qibin, 1986 Guanyintan Formation, China

Toarcian The Toarcian is, in the ICS' geologic timescale, an age and stage in the Early or Lower Jurassic. It spans the time between 182.7 Ma (million years ago) and 174.1 Ma. It follows the Pliensbachian and is followed by the Aalenian. The Toarcian ...

Aalenian The Aalenian () is a subdivision of the Middle Jurassic Epoch/Series of the geologic timescale that extends from about 174.1 Ma to about 170.3 Ma (million years ago). It was preceded by the Toarcian and succeeded by the Bajocian. Stratigraphic ...

* †'' Tetraphalerus mongolicus'' Ponomarenko, 1986 Gurvan-Eren Formation, Mongolia, Aptian * †'' Tetraphalerus notatus'' Ponomarenko, 1997 Dzun-Bain Formation, Mongolia, Aptian *'' Tetraphalerus wagneri'' Waterhouse, 1901, recent, South America

Formerly assigned species

* †''Tetraphalerus antiquus'' Ponomarenko, 1964 assigned to '' Allophalerus'' * †''Tetraphalerus bontsaganensis'' Ponomarenko, 1997 assigned to ''Allophalerus'' * †''Tetraphalerus brevicapitis'' Pononarenko & Martinez-Delclos, 2000 considered indeterminate by Kirejtshuk, 2020 * †''Tetraphalerus fentaiensis'' (Ren, 1995) assigned to '' Monticupes'' * †''Tetraphalerus grandis'' Ponomarenko, 1964 assigned to ''

Omma ''Omma'' is a genus of beetles in the family Ommatidae. ''Omma'' is an example of a living fossil. The oldest species known, ''O. liassicum'', lived during the final stage of the Triassic (Rhaetian), over 200 million years ago, though the placeme ...

'' * †''Tetraphalerus incertus'' Ponomarenko, 1969 assigned to ''Allophalerus'' * †''Tetraphalerus laetus'' Lin Qibin, 1976 assigned to ''Zygadenia'' * †''Tetraphalerus latus'' Tan et al. 2007 assigned to ''Allophalerus'' * †''Tetraphalerus lindae'' Jarzembowski et al. 2017 assigned to '' Bukhkalius'' * †''Tetraphalerus longicollis'' Ponomarenko, 1997 assigned to ''Omma'' * †''Tetraphalerus maximus'' Ponomarenko, 1968 assigned to ''Allophalerus'' * †''Tetraphalerus okhotensis'' Ponomarenko in Cromov et al., 1993 assigned to ''Allophalerus'' * †''Tetraphalerus oligocenicus'' (Crowson, 1962) assigned to '' Tetraphalerites'' * †''Tetraphalerus tenuipes'' Ponomarenko, 1964 assigned to ''Allophalerus'' * †''Tetraphalerus verrucosus'' Ponomarenko, 1966 assigned to ''Allophalerus'' * †''Tetraphalerus surrectus'' (Ren, 1995) assigned to ''Monticupes''

References

Ommatidae Archostemata genera {{Ommatidae-stub