''Tetracentron sinense'' is a

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

native to Asia and the sole living species in the genus ''

Tetracentron ''Tetracentron'' is a genus of flowering plant with a sole living species being '' Tetracentron sinense'' and several extinct species. It was formerly considered the sole genus in the family Tetracentraceae, though it is now included in the famil ...

''. It was formerly considered the sole species in the family

Tetracentraceae Trochodendraceae is the only family of flowering plants in the order Trochodendrales. It comprises two extant genera, each with a single species along with up to five additional extinct genera and a number of extinct species. The living specie ...

, though ''Tetracentron'' is now included in the family

Trochodendraceae Trochodendraceae is the only family of flowering plants in the order Trochodendrales. It comprises two extant genera, each with a single species along with up to five additional extinct genera and a number of extinct species. The living specie ...

together with the genus ''

Trochodendron ''Trochodendron'' is a genus of flowering plants with one living species, ''Trochodendron aralioides'', and six extinct species known from the fossil record. It was often considered the sole genus in the family Trochodendraceae, though botanists ...

''.

Range and habitat

It is native to southern

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

, northern

Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...

and the eastern

Himalaya The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...

(eastern

Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is mai ...

Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainous ...

Northeast India , native_name_lang = mni , settlement_type = , image_skyline = , image_alt = , image_caption = , motto = , image_map = Northeast india.png , ...

, and northern

Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John C. Wells, Joh ...

), where it grows at altitudes of along streams and forest margins in broad-leaved evergreen and mixed evergreen-deciduous forests.

Morphology

It is a

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

growing to tall. The

leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...

are

deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...

(the ''Flora of China'' reporting it as evergreen is an error), borne singly at the apex of short

spur shoot In botany, a plant shoot consists of any plant stem together with its appendages, leaves and lateral buds, flowering stems, and flower buds. The new growth from seed germination that grows upward is a shoot where leaves will develop. In the spri ...

s, each leaf dark green, broad heart-shaped, long and broad, with a rugose surface and a serrated margin. The spur shoots bear one leaf each year, slowly lengthening with each subsequent year. The

flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...

s are inconspicuous, yellowish green, without petals, produced on slender

catkin A catkin or ament is a slim, cylindrical flower cluster (a spike), with inconspicuous or no petals, usually wind-pollinated (anemophilous) but sometimes insect-pollinated (as in ''Salix''). They contain many, usually unisexual flowers, arranged cl ...

s long; each flower is diameter. The

fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in particu ...

is a follicle diameter, containing 4–6

seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...

s. ''Tetracentron'' and ''Trochodendron'' share the very unusual feature of lacking

vessel element A vessel element or vessel member (also called trachea or xylem vessel) is one of the cell types found in xylem, the water conducting tissue of plants. Vessel elements are typically found in angiosperms (flowering plants) but absent from most gy ...

s in the

wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin th ...

, something not typical in angiosperms. This has long been considered a very primitive character, resulting in the classification of these two genera in a basal position in the angiosperms; however,

molecular phylogenetics Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

research by the

Angiosperm Phylogeny Group The Angiosperm Phylogeny Group (APG) is an informal international group of systematic botanists who collaborate to establish a consensus on the taxonomy of flowering plants (angiosperms) that reflects new knowledge about plant relationships disco ...

and others has shown that these two genera are not

basal angiosperms The basal angiosperms are the flowering plants which diverged from the lineage leading to most flowering plants. In particular, the most basal angiosperms were called the ANITA grade, which is made up of '' Amborella'' (a single species of shrub ...

, but related basal eudicots. This suggests that the absence of vessel elements in this group is a secondarily evolved character, not a primitive one. image:Tetracentron sinense - Arbre-2.jpg, Branches image:Tetracentron sinense - Feuilles et fleurs-2.jpg, Leaves and flowers image:Tetracentron sinense - Feuilles et fleurs-3.jpg, Leaves and flowers image:Tetracentron sinense - Feuilles et fleurs.jpg, Leaves and flowers image:Tetracentron sinense - Couleur automnale.jpg, Fall color image:Tetracentron sinense - Jeunes graines-2.jpg, Young seeds image:Tetracentron sinense - Jeunes graines.jpg, Young seeds

References

External links

*Flora of China
''Tetracentron''
an
Tetracentraceae
{{Taxonbar, from=Q1418111 Trochodendrales Trees of Bhutan Trees of China Flora of East Himalaya Trees of Myanmar Trees of Nepal Trees of Vietnam Taxa named by Daniel Oliver