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''Plukenetia conophora'', also called Nigerian walnut, and conophore, is a climbing
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
in the genus '' Plukenetia''. It is not related to the
walnut A walnut is the edible seed of a drupe of any tree of the genus ''Juglans'' (family Juglandaceae), particularly the Persian or English walnut, '' Juglans regia''. Although culinarily considered a "nut" and used as such, it is not a true ...
, being so named because its nuts bear a superficial resemblance to the walnut. It is native to tropical western and central
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
from
Togo Togo (), officially the Togolese Republic (french: République togolaise), is a country in West Africa. It is bordered by Ghana to the west, Benin to the east and Burkina Faso to the north. It extends south to the Gulf of Guinea, where its c ...
to
Congo Congo or The Congo may refer to either of two countries that border the Congo River in central Africa: * Democratic Republic of the Congo, the larger country to the southeast, capital Kinshasa, formerly known as Zaire, sometimes referred to a ...
and in
Sierra Leone Sierra Leone,)]. officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered by Liberia to the southeast and Guinea surrounds the northern half of the nation. Covering a total area of , Sierra ...
. It is abundant in the
Nigeria Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf o ...
,
Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...
,
Republic of the Congo The Republic of the Congo (french: République du Congo, ln, Republíki ya Kongó), also known as Congo-Brazzaville, the Congo Republic or simply either Congo or the Congo, is a country located in the western coast of Central Africa to the w ...
and
Democratic Republic of Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...
. It prefers
rain-forest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainforest ...
hedge in half-shady places; low bush; secondary forest; plantations at elevations from . Although it is well recorded in Sierra Leone, it is apparently not indigenous to Sierra Leone, since it is not recorded in
Liberia Liberia (), officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to its north, Ivory Coast to its east, and the Atlantic Ocean ...
and
Ghana Ghana (; tw, Gaana, ee, Gana), officially the Republic of Ghana, is a country in West Africa. It abuts the Gulf of Guinea and the Atlantic Ocean to the south, sharing borders with Ivory Coast in the west, Burkina Faso in the north, and To ...
. Its presence in Sierra Leone is due to returning slaves for it is known to the Krio by its Yoruba (Nigerian) name. ''Plukenetia conophora'' is the only ''Plukenetia'' species native to
West Africa West Africa or Western Africa is the westernmost region of Africa. The United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Liberia, Mali, Maurit ...
or
Central Africa Central Africa is a subregion of the African continent comprising various countries according to different definitions. Angola, Burundi, the Central African Republic, Chad, the Democratic Republic of the Congo, the Republic of the Congo, ...
. Other ''Plukenetia'' species are indigenous to other parts of Africa, the
Indian Subcontinent The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...
,
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
, and
America The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
.


Description

It produces stems usually long, though they can be up to long. The seed is thin-shelled and about long. It is contained in a pod which may house;one shelled nut (single), two shelled nut (double) and three shelled nut. The walnut shells could be black or brown from the plant. The nut is whitish upon cracking from the shell. The nut has a thin layer in between two halves (when a nut is divided into two equal parts) of nut.


Taxonomy

''Plukenetia conophora'' belongs to the family Euphorbiaceae in the order Malpighiales of angiosperms in the plant kingdom. Euphorbiaceae is a large family of flowering plants with about 300 genera and 7,500 species. Members of the family are generally called spurge. This family occurs mainly in the tropics, with the majority of the species in the tropical America and Indo-Malayan regions. A large variety occurs in tropical Africa, but they are not as abundant or varied as in the two other tropical regions. However, Euphorbiaceae also has many species in nontropical areas such as the Mediterranean Basin, the Middle East, South Africa and southern USA. Of the three subfamilies of
Euphorbiaceae Euphorbiaceae, the spurge family, is a large family of flowering plants. In English, they are also commonly called euphorbias, which is also the name of a genus in the family. Most spurges, such as ''Euphorbia paralias'', are herbs, but some, e ...
Acalyphoideae The Acalyphoideae are a subfamily within the family Euphorbiaceae with 116 genera in 20 tribes. See also * Taxonomy of the Euphorbiaceae Here is a full taxonomy of the family Euphorbiaceae, according to the most recent molecular research. Thi ...
,
Crotonoideae The Crotonoideae (crotonoids) are a subfamily within the family Euphorbiaceae. See also * Taxonomy of the Euphorbiaceae Here is a full taxonomy of the family Euphorbiaceae, according to the most recent molecular research. This complex famil ...
and
Euphorbioideae The Euphorbioideae are a subfamily within the family Euphorbiaceae. See also * Taxonomy of the Euphorbiaceae Here is a full taxonomy of the family Euphorbiaceae, according to the most recent molecular research. This complex family previously ...
; only Acalyphoideae is native to Africa. The genus '' Plukenetia'' (
Euphorbiaceae Euphorbiaceae, the spurge family, is a large family of flowering plants. In English, they are also commonly called euphorbias, which is also the name of a genus in the family. Most spurges, such as ''Euphorbia paralias'', are herbs, but some, e ...
) is a
pantropical A pantropical ("all tropics") distribution is one which covers Tropics, tropical regions of both hemispheres. Examples of species include caecilians, modern sirenians and the plant genera ''Acacia'' and ''Bacopa''. ''Neotropical'' is a zoogeogra ...
genus of 19 species belonging to the tribe
Plukenetieae Plukenetieae is a tribe of plant of the family Euphorbiaceae. It comprises 3 subtribes and 14 genera. Tribe Plukenetieae (Benth.) Hutch. (Euphorbiaceae, Acalyphoideae) is a diverse pantropical lineage of ca. 17 genera and 350 species of twining ...
of the subfamily Acalyphoideae. Three sections or species groups of genus ''Plukenetia'' have been recognized. The two previously named genera, '' Tetracarpidium'' (synonym '' Angostylidium'') and '' Hedraiostylus'' (synonym '' Pterococcus''), are now treated as sections of Plukenetia. The monotypic ''Plukenetia'' sect. ''Tetracarpidium'' (''Angostylidium'') includes the African species ''Plukenetia conophora'', while ''Plukenetia'' sect. ''Hedraiostylus'' (''Pterococcus'') comprises two African species ('' Plukenetia africana'' and '' Plukenetia procumbens'') and one Asian species ('' Plukenetia corniculata''). A third species group restricted to Madagascar comprises three species; '' Plukenetia madagascariensis'', '' Plukenetia deciduas'', and '' Plukenetia ankaranensis''. However, ''Plukenetia conophora'' is still, in some literature, erroneously referred to by the genus ''Tetracarpidium''.


Uses


Food

''Plukenetia conophora'' is an important crop due to its multiple uses. ''Plukenetia conophora'' is widely cultivated for its nuts, which are cooked and consumed as snacks and often served with corn or rice. In West Africa, especially in Nigeria and Sierra Leone, as well as in Central African countries such as Cameroon, the seeds provide income to the rural people thereby improving their economy. Eaten boiled, it has a bitter aftertaste, usually observed upon drinking water. The seed can be ground into a powder and used with flour in making cakes. The nut, eaten raw, has a bitter taste comparable to that of
kola nut The term kola nut usually refers to the seeds of certain species of plant of the genus ''Cola'', placed formerly in the cocoa family Sterculiaceae and now usually subsumed in the mallow family Malvaceae (as subfamily Sterculioideae). These cola ...
. The nuts can also be roasted and eaten in the general diet, or added to cakes. The nuts are oil-bearing yielding 48–60% of a light golden colored oil with a taste resembling linseed oil. The oil is edible, although it not suitable for frying or soap manufacture, due to its quick drying property. The leaves are also edible and are often eaten with rice.


Traditional medicine

In Gabon, consumption of the seeds by husbands of pregnant women is believed to mitigate the risk of miscarriage. The fresh nuts are believed to be a snake antivenin.


Other uses

The oil has thus been found useful in the formulation of wood vanish, stand oil, vulcanized oil for leather and rubber substitute.


Local names

*Awusa, Asala in
Yoruba The Yoruba people (, , ) are a West African ethnic group that mainly inhabit parts of Nigeria, Benin, and Togo. The areas of these countries primarily inhabited by Yoruba are often collectively referred to as Yorubaland. The Yoruba constitute ...
,
Nigeria Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf o ...
*Ukpa in
Igbo Igbo may refer to: * Igbo people, an ethnic group of Nigeria * Igbo language, their language * anything related to Igboland, a cultural region in Nigeria See also * Ibo (disambiguation) * Igbo mythology * Igbo music * Igbo art * * Igbo-Ukwu, a t ...
,
Nigeria Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf o ...
*Kaso, Ngak in
Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...


References

{{Taxonbar, from=Q15371019 Plukenetieae