Tessema Sensilis
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''Tessema sensilis'' is a little-known
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
, the only member of
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Tessema''. It belongs to the grass moth
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
(Crambidae), and therein to the large
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Spilomelinae; at the time of its description, these were still included in subfamily
Pyraustinae Pyraustinae is a large subfamily of the lepidopteran family Crambidae, the crambid snout moths. It currently includes over 1,400 species; most of them tropical but some found in temperate regions including both North America and Europe. The Py ...
and the entire Crambidae was then merged with the snout moths (family Pyralidae). While its detailed relationships are undetermined, it may be a close relative of ''
Herpetogramma ''Herpetogramma'' is a genus of moths of the family Crambidae described by Julius Lederer in 1863. It currently comprises 106 species and is found in North America, Eurasia, Australia, New Zealand, Central and South America. Of the few specie ...
'' and/or '' Pilocrocis''.Clarke (1986) Though there is no particular reason to assume it is very rare, this moth is inconspicuous and has in fact only been recorded a single time so far, on January 23, 1968; it took almost 20 years to realize it was a new and distinct animal. It inhabits the island of Nuku Hiva, in the
Marquesas Islands The Marquesas Islands (; french: Îles Marquises or ' or '; Marquesan: ' ( North Marquesan) and ' ( South Marquesan), both meaning "the land of men") are a group of volcanic islands in French Polynesia, an overseas collectivity of France in th ...
of
Polynesia Polynesia () "many" and νῆσος () "island"), to, Polinisia; mi, Porinihia; haw, Polenekia; fj, Polinisia; sm, Polenisia; rar, Porinetia; ty, Pōrīnetia; tvl, Polenisia; tkl, Polenihia (, ) is a subregion of Oceania, made up of ...
, where it may be
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
and only specimen (
USNM The National Museum of Natural History is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. It has free admission and is open 364 days a year. In 2021, with 7. ...
100735, genitalia on
microscopic slide A microscope slide is a thin flat piece of glass, typically 75 by 26 mm (3 by 1 inches) and about 1 mm thick, used to hold objects for examination under a microscope. Typically the object is mounted (secured) on the slide, and then b ...
USNM 25220) was collected on Tunoa Ridge about 885 m (2900 ft)
ASL American Sign Language (ASL) is a natural language that serves as the predominant sign language of Deaf communities in the United States of America and most of Anglophone Canada. ASL is a complete and organized visual language that is express ...
, near the scenic lookout at .


Description and ecology

''T. sensilis'' is a smallish smooth-bodied and notably long-legged moth, mid-sized by grass moth standards with a wingspan of 39 mm in the only known specimen. It is mostly a medium yellowish-brown in color, and in the details closely resembles '' Herpetogramma fimbrialis'', and somewhat less so such species as '' Palpita cupripennalis''"''Glyphodes cypripennalis''" in Clarke (1986) is a ''
lapsus In philology, a lapsus (Latin for "lapse, slip, error") is an involuntary mistake made while writing or speaking. Investigations In 1895 an investigation into verbal slips was undertaken by a philologist and a psychologist, Rudolf Meringer and K ...
''
and '' Glyphodes argyritis''. The head is slightly scaly, with a well-developed
proboscis A proboscis () is an elongated appendage from the head of an animal, either a vertebrate or an invertebrate. In invertebrates, the term usually refers to tubular mouthparts used for feeding and sucking. In vertebrates, a proboscis is an elong ...
and squamiform
labial The term ''labial'' originates from '' Labium'' (Latin for "lip"), and is the adjective that describes anything of or related to lips, such as lip-like structures. Thus, it may refer to: * the lips ** In linguistics, a labial consonant ** In zoolog ...
palps, which are white on the underside; the small
maxillary palp Maxillary means "related to the maxilla (upper jaw bone)". Terms containing "maxillary" include: *Maxillary artery *Maxillary nerve *Maxillary prominence *Maxillary sinus {{disambig ...
s are simple knobs which project forward. Its greyish
antenna Antenna ( antennas or antennae) may refer to: Science and engineering * Antenna (radio), also known as an aerial, a transducer designed to transmit or receive electromagnetic (e.g., TV or radio) waves * Antennae Galaxies, the name of two collid ...
e have brownish spots at the base and – at least in the male – are long (almost 2 cm each) and hairy. Between the antenna bases and the compound eyes there is a small crescent of white scales on each side. The tegula has buff scales at the hind end. The
abdomen The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso. ...
is reddish-grey, but the 8th segment is white in the front part, reddish in the hind part. The end of the abdomen bears a tuft of elongated scales which are buff with reddish tips above, and entirely reddish below. The underside of the body is white. The
legs A leg is a weight-bearing and locomotive anatomical structure, usually having a columnar shape. During locomotion, legs function as "extensible struts". The combination of movements at all joints can be modeled as a single, linear element ...
are white at the base, shading to brownish on the femur (forelegs) or tibia (mid- and hindlegs). The straight-margined forewings have a somewhat drawn-out but blunt tip and 12 veins. Of the latter, lb and 2 are single, 3-5 approach at their base, 5-7 run somewhat parallel and not far apart from each other, 8 and 9 have a long stalk leading to wingtip or leading edge, respectively; the 10th vein
anastomoses An anastomosis (, plural anastomoses) is a connection or opening between two things (especially cavities or passages) that are normally diverging or branching, such as between blood vessels, leaf veins, or streams. Such a connection may be normal ...
with the stalks of the preceding two, the 11th attaches to the outer fourth of the wing cell, and the 12th from its base. The hindwings are somewhat angular too, and have 8 veins. Of these, the first two run singly, while 3-5 approach, 6 and 7 join, and 7 and 8 anastomose at their bases. The only mark on the yellowish-brown wings is a short reddish transverse dash at the outer end of the forewing cell. Altogether, the most conspicuous differences from ''H. fimbrialis'' are the markedly shorter legs and antennae of the latter, as well as its white abdominal tuft. In addition, ''H. fimbrialis'' has upturned (not squamiform) labial palps and stalked (not anastomosing) hindwing veins 7 and 8. The male genitalia of ''T. sensilis'' are symmetrical and overall only lightly sclerotized (hardened) except for the stout
aedeagus An aedeagus (plural aedeagi) is a reproductive organ of male arthropods through which they secrete sperm from the testes during copulation with a female. It can be thought of as the insect equivalent of a mammal's penis, though the comparison ...
, which has a heavily sclerotized rod on the underside. Gnathos and socius are absent. The
uncus The uncus is an anterior extremity of the parahippocampal gyrus. It is separated from the apex of the temporal lobe by a slight fissure called the incisura temporalis (also called rhinal sulcus). Although superficially continuous with the hipp ...
is slender, curved, dilated, and covered in bristles at the tip. The clasper's harpe is barely sclerotized, forming a ridge on the costa, a narrow stripe on the sacculus, and a curved process emerging from the harpe's center; the cucullus is broadly rounded. The vinculum is broad and truncated, with a bulge in the middle; the tegumen is arched, and the
anellus ''Anellus'' is a monotypic moth genus of the family Erebidae. Its only species, ''Anellus edai'', is known from Ishigaki Island, which is southwest of Japan. Both the genus and the species were first described by Michael Fibiger in 2008. Adults ...
forms a small almost triangular plate with light sclerotized flanges at the hind sides. The female genitalia are of course still unknown. Ecologically, essentially nothing is known about this species. Its
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
is likely light woodland rich in
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s,
fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...
s,
moss Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hor ...
es and
lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Bidens henryi ''Bidens'' is a genus of flowering plants in the aster family, Asteraceae.''Bidens''.
Flo ...
'', ''
Cheirodendron bastardianum ''Cheirodendron'' is a genus of flowering plant in the family Araliaceae. All six species in the genus are Endemism, endemic to Polynesia. The five Hawaiian species are generally called ''ōlapa'', and occur in Hawaiian tropical rainforests#Wet ...
'', ''
Glochidion ramiflorum ''Glochidion'' is a genus of flowering plants, of the family (biology), family Phyllanthaceae, known as cheese trees or buttonwood in Australia, and leafflower trees in the scientific literature. It comprises about 300 species, distributed from M ...
'', ''
Metrosideros collina ''Metrosideros collina'' is a species of flowering plant in the family Myrtaceae. It is native to French Polynesia and the Cook Islands. Taxonomy The species was first formally described by botanist Johann Reinhold Forster and his son Georg F ...
'', ''
Pandanus ''Pandanus'' is a genus of monocots with some 750 accepted species. They are palm-like, dioecious trees and shrubs native to the Old World tropics and subtropics. The greatest number of species are found in Madagascar and Malaysia. Common names ...
'', and ''
Vaccinium cereum ''Vaccinium'' is a common and widespread genus of shrubs or dwarf shrubs in the heath family (Ericaceae). The fruits of many species are eaten by humans and some are of commercial importance, including the cranberry, blueberry, bilberry (who ...
''.


Footnotes


References

* (1986): Pyralidae and Microlepidoptera of the Marquesas Archipelago. ''Smithsonian Contributions to Zoology'' 416: 1-485
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External links


Global Pyraloidea database
{{Taxonbar, from1=Q7705654, from2=Q17272380 Spilomelinae Fauna of the Marquesas Islands