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A tepui , or tepuy (), is a table-top mountain or mesa found in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, especially in
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
and western
Guyana Guyana ( or ), officially the Cooperative Republic of Guyana, is a country on the northern mainland of South America. Guyana is an indigenous word which means "Land of Many Waters". The capital city is Georgetown. Guyana is bordered by the ...
. The word tepui means "house of the gods" in the native tongue of the Pemon, the
indigenous people Indigenous peoples are culturally distinct ethnic groups whose members are directly descended from the earliest known inhabitants of a particular geographic region and, to some extent, maintain the language and culture of those original people ...
who inhabit the Gran Sabana. Tepuis tend to be found as isolated entities rather than in connected ranges, which makes them the host of a unique array of
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
plant and animal species. Some of the most outstanding tepuis are Auyantepui, Autana,
Neblina Neblina is a heavy metal band from Angola, founded in 2001. They released an album in 2006, ''Innocence Falls in Decay''. In the same year, they played a number of shows in Germany, for the 2006 FIFA World Cup. In 2007, they performed at Windh ...
, and
Mount Roraima Mount Roraima ( es, Monte Roraima; Tepuy Roraima; Cerro Roraima pt, Monte Roraima ) is the highest of the Pakaraima chain of tepuis (table-top mountain) or plateaux in South America. It is located at the junction of Venezuela, Brazil and Guyana. ...
. They are typically composed of sheer blocks of
Precambrian The Precambrian (or Pre-Cambrian, sometimes abbreviated pꞒ, or Cryptozoic) is the earliest part of Earth's history, set before the current Phanerozoic Eon. The Precambrian is so named because it preceded the Cambrian, the first period of the ...
quartz arenite A quartz arenite or quartzarenite is a sandstone composed of greater than 90% detrital quartz. Quartz arenites are the most mature sedimentary rocks possible, and are often referred to as ultra- or super-mature, and are usually cemented by sili ...
sandstone Sandstone is a clastic sedimentary rock composed mainly of sand-sized (0.0625 to 2 mm) silicate grains. Sandstones comprise about 20–25% of all sedimentary rocks. Most sandstone is composed of quartz or feldspar (both silicates) ...
that rise abruptly from the jungle, giving rise to spectacular natural scenery. Auyantepui is the source of Angel Falls, the world's tallest
waterfall A waterfall is a point in a river or stream where water flows over a vertical drop or a series of steep drops. Waterfalls also occur where meltwater drops over the edge of a tabular iceberg or ice shelf. Waterfalls can be formed in several wa ...
.


Morphology

These table-top mountains are the remains of a large sandstone plateau that once covered the granite basement complex between the north border of the
Amazon Basin The Amazon basin is the part of South America drained by the Amazon River and its tributaries. The Amazon drainage basin covers an area of about , or about 35.5 percent of the South American continent. It is located in the countries of Bolivi ...
and the
Orinoco The Orinoco () is one of the longest rivers in South America at . Its drainage basin, sometimes known as the Orinoquia, covers , with 76.3 percent of it in Venezuela and the remainder in Colombia. It is the fourth largest river in the wor ...
, between the Atlantic coast and the Rio Negro. This area is part of the remnants of the supercontinent
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
. Throughout the course of the
history of Earth The history of Earth concerns the development of planet Earth from its formation to the present day. Nearly all branches of natural science have contributed to understanding of the main events of Earth's past, characterized by constant geologi ...
, the plateau was eroded, and the tepuis were formed from the remaining
monadnock An inselberg or monadnock () is an isolated rock hill, knob, ridge, or small mountain that rises abruptly from a gently sloping or virtually level surrounding plain. In Southern Africa a similar formation of granite is known as a koppie, a ...
s. There are 115 such mesas in the Gran Sabana in the south-east of Venezuela on the border with
Guyana Guyana ( or ), officially the Cooperative Republic of Guyana, is a country on the northern mainland of South America. Guyana is an indigenous word which means "Land of Many Waters". The capital city is Georgetown. Guyana is bordered by the ...
and
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
, where the highest concentration of tepuis is found. The precipitous
mountain A mountain is an elevated portion of the Earth's crust, generally with steep sides that show significant exposed bedrock. Although definitions vary, a mountain may differ from a plateau in having a limited Summit (topography), summit area, and ...
s tower over the surrounding area by up to . Tepuis range in elevation from . The total surface area of all 115 tepuis is approximately . Because of their great age, some tepuis exhibit surface features and caves typical of
karst Karst is a topography formed from the dissolution of soluble rocks such as limestone, dolomite, and gypsum. It is characterized by underground drainage systems with sinkholes and caves. It has also been documented for more weathering-resistant ro ...
topography, formed in more water-soluble rocks such as
limestone Limestone ( calcium carbonate ) is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of . Limestone forms whe ...
. Caves here include the
Abismo Guy Collet The Abismo Guy Collet (SP-090) is the deepest cave in South America, and the deepest in the world that is formed in quartzite. Located in a tepui in northern Brazil near the border with Venezuela, this cave was first explored by cavers in 2006 to ...
, the deepest quartzite cave in the world. Some of the
mesas A mesa is an isolated, flat-topped elevation, ridge or hill, which is bounded from all sides by steep escarpments and stands distinctly above a surrounding plain. Mesas characteristically consist of flat-lying soft sedimentary rocks capped by ...
are pocked with giant
sinkhole A sinkhole is a depression or hole in the ground caused by some form of collapse of the surface layer. The term is sometimes used to refer to doline, enclosed depressions that are locally also known as ''vrtače'' and shakeholes, and to openi ...
s up to in diameter and with sheer walls up to deep. These sinkholes are formed when the roofs of tunnels carved by underground rivers collapse.


Flora and fauna

The plateaus of the tepuis are completely isolated from the ground forest, making them ecological islands.Zimmer, Carl
It’s Not So Lonely at the Top: Ecosystems Thrive High in the Sky
The New York Times ''The New York Times'' (''the Times'', ''NYT'', or the Gray Lady) is a daily newspaper based in New York City with a worldwide readership reported in 2020 to comprise a declining 840,000 paid print subscribers, and a growing 6 million paid ...
website, May 7, 2012, and published in the New York edition, p. D3 on May 8, 2012.
The altitude causes them to have a different climate from the ground forest. The top presents cool temperatures with frequent rainfall, while the bases of the mountains have a tropical, warm and humid climate. The isolation has led to the presence of
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
flora and fauna through evolution over millennia of a different world of animals and plants, cut off from the rest of the world by the imposing rock walls. Some tepui sinkholes contain species that have evolved in these "islands within islands" that are unique to that sinkhole. The tepuis are often referred to as the
Galápagos Islands The Galápagos Islands (Spanish: , , ) are an archipelago of volcanic islands. They are distributed on each side of the equator in the Pacific Ocean, surrounding the centre of the Western Hemisphere, and are part of the Republic of Ecuador ...
of the mainland, having a large number of unique plants and animals not found anywhere else in the world. The floors of the mesas are poor in nutrients, which has led to a rich variety of carnivorous plants, such as ''
Drosera ''Drosera'', which is commonly known as the sundews, is one of the largest genus, genera of carnivorous plants, with at least 194 species. 2 volumes. These members of the family Droseraceae lure, capture, and digest insects using stalked mucil ...
'' and most species of ''
Heliamphora The genus ''Heliamphora'' ( or ; Greek: ''helos'' "marsh" and ''amphoreus'' " amphora") contains 23 species of pitcher plants endemic to South America.McPherson, S., A. Wistuba, A. Fleischmann & J. Nerz 2011. '' Sarraceniaceae of South America''. ...
'', as well as a wide variety of orchids and bromeliads. The weathered, craggy nature of the rocky ground means no layers of
humus In classical soil science, humus is the dark organic matter in soil that is formed by the decomposition of plant and animal matter. It is a kind of soil organic matter. It is rich in nutrients and retains moisture in the soil. Humus is the Lati ...
are formed. It has been hypothesized that endemics on tepuis represent relict fauna and flora that underwent
vicariant speciation Allopatric speciation () – also referred to as geographic speciation, vicariant speciation, or its earlier name the dumbbell model – is a mode of speciation that occurs when biological populations become geographically isolated from ...
when the plateau became fragmented over geological time. However, recent studies suggest that tepuis are not as isolated as originally believed. For example, an endemic group of treefrogs, ''
Tepuihyla ''Tepuihyla'', commonly known as Amazon tree frogs or Tepui tree frogs, is a genus of frogs in the family Hylidae found in mountains of eastern and south-eastern Venezuela and Guyana, and likely in adjacent Brazil. A tepui is a table-top mountai ...
'', have diverged after the tepuis were formed; that is, speciation followed colonization from the lowlands. The tepuis, also known as 'islands above the rainforest', are a challenge for researchers, as they are home to a high number of new species that have yet to be described. A few of these mountains are cloaked by thick clouds for nearly the entire year. Their surfaces could previously only be photographed by helicopter radar equipment. Many tepuis are in the
Canaima National Park Canaima National Park ( es, Parque Nacional Canaima) is a park in south-eastern Venezuela that roughly occupies the same area as the Gran Sabana region. It is located in Bolívar State, reaching the borders with Brazil and Guyana. History Ca ...
in Venezuela, which has been classified as a
World Heritage Site A World Heritage Site is a landmark or area with legal protection by an international convention administered by the United Nations Educational, Scientific and Cultural Organization (UNESCO). World Heritage Sites are designated by UNESCO for h ...
by
UNESCO The United Nations Educational, Scientific and Cultural Organization is a specialized agency of the United Nations (UN) aimed at promoting world peace and security through international cooperation in education, arts, sciences and culture. It ...
.


Selected tepuis

A few of the most notable of the 60 tepuis: * Auyantepui is the largest of the tepuis with a surface area of . Angel Falls, the highest waterfall in the world, drops from a cleft in the summit. *
Mount Roraima Mount Roraima ( es, Monte Roraima; Tepuy Roraima; Cerro Roraima pt, Monte Roraima ) is the highest of the Pakaraima chain of tepuis (table-top mountain) or plateaux in South America. It is located at the junction of Venezuela, Brazil and Guyana. ...
, also known as Roraima Tepui. A report by the noted South American researcher Robert Schomburgk inspired the Scottish author
Arthur Conan Doyle Sir Arthur Ignatius Conan Doyle (22 May 1859 – 7 July 1930) was a British writer and physician. He created the character Sherlock Holmes in 1887 for ''A Study in Scarlet'', the first of four novels and fifty-six short stories about Ho ...
to write his novel ''
The Lost World The lost world is a subgenre of the fantasy or science fiction genres that involves the discovery of an unknown Earth civilization. It began as a subgenre of the late- Victorian adventure romance and remains popular into the 21st century. The g ...
'' about the discovery of a living prehistoric world full of dinosaurs and other primordial creatures. The borders of
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
,
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
, and
Guyana Guyana ( or ), officially the Cooperative Republic of Guyana, is a country on the northern mainland of South America. Guyana is an indigenous word which means "Land of Many Waters". The capital city is Georgetown. Guyana is bordered by the ...
meet on the top. * Matawi Tepui, also known as Kukenán, because it is the source of the Kukenán River, is considered the "place of the dead" by the local Pemon Peoples. Located next to Mount Roraima in Venezuela. * Autana Tepui stands above the forest floor. A unique cave runs from one side of the mountain to the other. * Sarisariñama Tepui, famous for its almost perfectly circular sinkholes that go straight down from the mountain top – the largest such sinkhole is in diameter and depth (purportedly created by groundwater erosion). They harbour an
ecosystem An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syste ...
composed of unique plant and animal species at the bottom. * Ilú-Tramen Massif is the most northerly mountain in the chain that stretches along the Venezuelan-Guyana border from Roraima in the south.


Popular culture

The hypothesis that endemic fauna and flora of tepuis represent remnants of ancient species was an inspiration to
Arthur Conan Doyle Sir Arthur Ignatius Conan Doyle (22 May 1859 – 7 July 1930) was a British writer and physician. He created the character Sherlock Holmes in 1887 for ''A Study in Scarlet'', the first of four novels and fifty-six short stories about Ho ...
's novel ''
The Lost World The lost world is a subgenre of the fantasy or science fiction genres that involves the discovery of an unknown Earth civilization. It began as a subgenre of the late- Victorian adventure romance and remains popular into the 21st century. The g ...
'' (1912), which was set on a tepui. In the 1990 film ''
Arachnophobia Arachnophobia is a specific phobia brought about by the irrational fear of spiders and other arachnids such as scorpions. Signs and symptoms People with arachnophobia tend to feel uneasy in any area they believe could harbour spiders or that h ...
'', a new species of spider with a highly potent, deadly venom that lives in social colonies and is the implied apex predator of its environment is discovered in a sinkhole enclosed by a tepui in the Venezuelan rainforest, also alluding to the unique ecological conditions of tepuis. Much of the story of 2009 Pixar
Disney The Walt Disney Company, commonly known as Disney (), is an American multinational mass media and entertainment conglomerate headquartered at the Walt Disney Studios complex in Burbank, California. Disney was originally founded on October ...
film '' Up'' takes place among the tepuis. The film also includes depictions of the numerous rock formations and an Angel Falls-like waterfall called "Paradise Falls". In a scene near the beginning of Disney's "
Dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
", a pteranodon flies over tepuis.
Steve Backshall Stephen James Backshall (born 21 April 1973) is an English naturalist, explorer, presenter and writer, best known for BBC TV's ''Deadly 60''. His other BBC work includes being part of the expedition teams in ''Lost Land of the Tiger'', ''Lo ...
was part of the first expedition to successfully climb
Mount Upuigma Upuigma-tepui, also known as El Castillo, is a tepui in Bolívar state, Venezuela.Huber, O. (1995). Geographical and physical features. In: P.E. Berry, B.K. Holst & K. Yatskievych (eds.) '' Flora of the Venezuelan Guayana. Volume 1. Introduction. ...
."Conquering a virgin"
''The Times'', 11 November 2007
The expedition was part of the program ''
Lost Land of the Jaguar ''Lost Land of the Jaguar'' is a 2008 British nature documentary series on the fauna of Guyana's rainforest. The four presenters are George McGavin, Steve Backshall, Justine Evans, and Gordon Buchanan. The series is a production of the BBC Nat ...
'' on
BBC One BBC One is a British free-to-air public broadcast television network owned and operated by the BBC. It is the corporation's flagship network and is known for broadcasting mainstream programming, which includes BBC News television bulletins, p ...
in 2008. On the summit they discovered an endemic species of frog and mouse, and also footprints of an unidentified mammal. In the novel ''The 6th Extinction'' (2014) by James Rollins, a tepui in the north of Brazil was featured as a secret lair for the villain Dr. Cutter Elwes.


See also

* * *


References

;Citations ;Bibliography * Much of the text of this article comes from the corresponding German-language Wikipedia article, retrieved on 16 February 2006, which uses the following sources: ** Uwe George: ''Inseln in der Zeit''. GEO - Gruner + Jahr AG & Co., Hamburg, . ** Roland Stuckardt: ''Sitze der Götter''. terra - Heft 3/2004,
Tecklenborg Verlag Tecklenborg Verlag is a publishing house based in Steinfurt, North Rhine-Westphalia, Germany that is internationally recognized for its illustrated books on wild animals, and for their nature-related magazines, which include ''Naturfoto'' and '' ...
, Steinfurt. * National Geographic Magazine, May 1989, "Venezuela's Islands in Time," pp. 526–561


External links


The Making of the Tepuis Film : "The Living Edens : The Lost World"

Mongabay.com
– pictures from Tepuis in Venezuela. {{Tepuis, state=expanded . . . . Erosion landforms Mesas Tables (landform) Oronyms Natural history of South America Flora of northern South America Ecoregions of South America Ecoregions of Brazil Ecoregions of Guyana Ecoregions of Suriname Ecoregions of Venezuela Neotropical tropical and subtropical moist broadleaf forests Montane forests