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Temperate broadleaf and mixed forest is a
temperate climate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
terrestrial
habitat type In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physica ...
defined by the
World Wide Fund for Nature The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the Wor ...
, with
broadleaf tree A broad-leaved, broad-leaf, or broadleaf tree is any tree within the diverse botanical group of angiosperms that has flat leaves and produces seeds inside of fruits. It is one of two general types of trees, the other being a conifer, a tree with n ...
ecoregion An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas of ...
s, and with conifer and broadleaf tree
mixed coniferous forest Mixed coniferous forest is a vegetation type dominated by a mixture of broadleaf trees and conifers.Sierra Nevada Wildflowers, Karen Wiese, 2nd ed, 2013, p 13–14 It is generally located in mountains, below the upper montane vegetation type. Sie ...
ecoregions. These forests are richest and most distinctive in central China and eastern North America, with some other globally distinctive ecoregions in the
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia (country), Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range ...
, the
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 10 ...
,
Southern Europe Southern Europe is the southern region of Europe. It is also known as Mediterranean Europe, as its geography is essentially marked by the Mediterranean Sea. Definitions of Southern Europe include some or all of these countries and regions: Alba ...
,
Australasia Australasia is a region that comprises Australia, New Zealand and some neighbouring islands in the Pacific Ocean. The term is used in a number of different contexts, including geopolitically, physiogeographically, philologically, and ecologi ...
, Southwestern
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
and the
Russian Far East The Russian Far East (russian: Дальний Восток России, r=Dal'niy Vostok Rossii, p=ˈdalʲnʲɪj vɐˈstok rɐˈsʲiɪ) is a region in Northeast Asia. It is the easternmost part of Russia and the Asian continent; and is admin ...
.


Ecology

The typical structure of these forests includes four layers. * The uppermost layer is the
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an ...
composed of tall mature trees ranging from high. Below the canopy is the three-layered,
shade-tolerant In ecology, shade tolerance is a plant's ability to tolerate low light levels. The term is also used in horticulture and landscaping, although in this context its use is sometimes imprecise, especially in labeling of plants for sale in commercial ...
understory In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but abo ...
that is roughly shorter than the canopy. * The top layer of the understory is the sub-canopy composed of smaller mature trees,
sapling In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s, and suppressed juvenile canopy layer trees awaiting an opening in the canopy. * Below the sub-canopy is the
shrub layer Stratification in the field of ecology refers to the vertical layering of a habitat; the arrangement of vegetation in layers. It classifies the layers (sing. ''stratum'', pl. ''strata'') of vegetation largely according to the different heights to w ...
, composed of low growing woody plants. * Typically the lowest growing (and most diverse) layer is the
ground cover Groundcover or ground cover is any plant that grows over an area of ground. Groundcover provides protection of the topsoil from erosion and drought. In an ecosystem, the ground cover forms the layer of vegetation below the shrub layer known as ...
or
herbaceous layer Stratification in the field of ecology refers to the vertical layering of a habitat; the arrangement of vegetation in layers. It classifies the layers (sing. ''stratum'', pl. ''strata'') of vegetation largely according to the different heights to w ...
.


Trees

In the Northern hemisphere, characteristic dominant
broadleaf tree A broad-leaved, broad-leaf, or broadleaf tree is any tree within the diverse botanical group of angiosperms that has flat leaves and produces seeds inside of fruits. It is one of two general types of trees, the other being a conifer, a tree with n ...
s in this biome include oaks (''
Quercus An oak is a tree or shrub in the genus ''Quercus'' (; Latin "oak tree") of the beech family, Fagaceae. There are approximately 500 extant species of oaks. The common name "oak" also appears in the names of species in related genera, notably ...
'' spp.), beeches (''
Fagus Beech (''Fagus'') is a genus of deciduous trees in the family Fagaceae, native to temperate Europe, Asia, and North America. Recent classifications recognize 10 to 13 species in two distinct subgenera, ''Engleriana'' and ''Fagus''. The ''Engle ...
'' spp.), maples ('' Acer'' spp.), or birches (''
Betula A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech- oak family Fagaceae. The genus ''Betula'' contains 30 t ...
'' spp.). The term "mixed forest" comes from the inclusion of coniferous trees as a canopy component of some of these forests. Typical coniferous trees include pines (''
Pinus A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanical Garden ...
'' spp.), firs (''
Abies Firs (''Abies'') are a genus of 48–56 species of evergreen coniferous trees in the family Pinaceae. They are found on mountains throughout much of North and Central America, Europe, Asia, and North Africa. The genus is most closely related t ...
'' spp.), and spruces ('' Picea'' spp.). In some areas of this biome, the conifers may be a more important canopy species than the broadleaf species. In the Southern Hemisphere, endemic genera such as ''
Nothofagus ''Nothofagus'', also known as the southern beeches, is a genus of 43 species of trees and shrubs native to the Southern Hemisphere in southern South America (Chile, Argentina) and Australasia (east and southeast Australia, New Zealand, New Gui ...
'' and ''
Eucalyptus ''Eucalyptus'' () is a genus of over seven hundred species of flowering trees, shrubs or mallees in the myrtle family, Myrtaceae. Along with several other genera in the tribe Eucalypteae, including '' Corymbia'', they are commonly known as e ...
'' occupy this biome, and most coniferous trees (members of the
Araucariaceae Araucariaceae – also known as araucarians – is an extremely ancient family of coniferous trees. The family achieved its maximum diversity during the Jurassic and Cretaceous periods and the early Cenozoic, when it was distributed almost worldw ...
and
Podocarpaceae Podocarpaceae is a large family of mainly Southern Hemisphere conifers, known in English as podocarps, comprising about 156 species of evergreen trees and shrubs.James E. Eckenwalder. 2009. ''Conifers of the World''. Portland, Oregon: Timber Pr ...
) occur in mixtures with broadleaf species, and are classed as broadleaf and mixed forests.


Climate

Temperate broadleaf and mixed forests occur in areas with distinct warm and cool seasons that give them moderate annual average temperatures — . These forests occur in relatively warm and rainy climates, sometimes also with a distinct
dry season The dry season is a yearly period of low rainfall, especially in the tropics. The weather in the tropics is dominated by the tropical rain belt, which moves from the northern to the southern tropics and back over the course of the year. The te ...
. A dry season occurs in the winter in East Asia and in summer on the wet fringe of the
Mediterranean climate A Mediterranean climate (also called a dry summer temperate climate ''Cs'') is a temperate climate sub-type, generally characterized by warm, dry summers and mild, fairly wet winters; these weather conditions are typically experienced in the ...
zones. Other areas, such as central eastern North America, have a fairly even distribution of rainfall; annual rainfall is typically over and often over , though it can go as low as in some parts of the Middle East and close to in the mountains of New Zealand and the Azores. Temperatures are typically moderate except in parts of Asia such as
Ussuriland Primorsky Krai (russian: Приморский край, r=Primorsky kray, p=prʲɪˈmorskʲɪj kraj), informally known as Primorye (, ), is a federal subject (a krai) of Russia, located in the Far East region of the country and is a part of the ...
, where temperate forests can occur despite very harsh conditions with very cold winters. The climates are typically humid for much of the year, usually appearing in the humid subtropical climate without summer being extremely hot and winter very mild, and in the
humid continental climate A humid continental climate is a climatic region defined by Russo-German climatologist Wladimir Köppen in 1900, typified by four distinct seasons and large seasonal temperature differences, with warm to hot (and often humid) summers and freezing ...
zones to the south of
tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless mou ...
and the generally subarctic
taiga Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruc ...
. In the
Köppen climate classification The Köppen climate classification is one of the most widely used climate classification systems. It was first published by German-Russian climatologist Wladimir Köppen (1846–1940) in 1884, with several later modifications by Köppen, notabl ...
they are represented respectively by '' Cfa'', '' Dfa/Dfb'' southern range and '' Cfb'', and more rarely, '' Csb'', ''
BSk BSK is a three-letter abbreviation that may refer to: Sports * OFK Beograd, Serbian football club * FK BSK Borča, Serbian football club * FK BSK Batajnica, Serbian football club * FK BSK Banja Luka, Bosnian-Herzegovinian football club * FK BSK L ...
'' and ''
Csa CSA may refer to: Arts and media * Canadian Screen Awards, annual awards given by the Academy of Canadian Cinema & Television * Commission on Superhuman Activities, a fictional American government agency in Marvel Comics * Crime Syndicate of Amer ...
''.


Ecoregions


Australasia


Eurasia


Americas


See also

*
Mixed coniferous forest Mixed coniferous forest is a vegetation type dominated by a mixture of broadleaf trees and conifers.Sierra Nevada Wildflowers, Karen Wiese, 2nd ed, 2013, p 13–14 It is generally located in mountains, below the upper montane vegetation type. Sie ...
* Kuchler plant association system * Mediterranean forests, woodlands, and scrub *
Temperate deciduous forest Temperate deciduous or temperate broad-leaf forests are a variety of temperate forest 'dominated' by trees that lose their leaves each year. They are found in areas with warm moist summers and cool winters. The six major areas of this forest type ...
*
Trees of the world A listing of lists of trees. * List of individual trees, including actual and mythical trees * List of largest giant sequoias * List of old growth forests * List of oldest trees * List of superlative trees **List of superlative trees in Sweden * ...


References


External links


World Wildlife Fund−WWF Biomes: Temperate broadleaf and mixed forests biome


* ttps://web.archive.org/web/20070310230140/http://forest.terraformers.ca/ Terraformers Canadian Forest Conservation Foundation {{Biomes Forests Terrestrial biomes