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Teleosauridae is a
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
typically marine
crocodylomorphs Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. During Mesozoic and early Cenozoic times, cr ...
similar to the modern
gharial The gharial (''Gavialis gangeticus''), also known as gavial or fish-eating crocodile, is a crocodilian in the family Gavialidae and among the longest of all living crocodilians. Mature females are long, and males . Adult males have a distinct b ...
that lived during the
Jurassic period The Jurassic ( ) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of ...
. Teleosaurids were
thalattosuchians Thalattosuchia is a clade of marine crocodylomorphs from the Early Jurassic to the Early Cretaceous that had a cosmopolitan distribution. They are colloquially referred to as marine crocodiles or sea crocodiles, though they are not members of Cro ...
closely related to the fully aquatic metriorhynchoids, but were less adapted to an open-ocean,
pelagic The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or wa ...
lifestyle. The family was originally coined to include all the semi-aquatic (i.e. non-metriorhynchoid) thalattosuchians and was equivalent to the modern
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
Teleosauroidea Teleosauroidea is an extinct superfamily of thalattosuchian crocodyliforms living from the Early Jurassic to the Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier o ...
. However, as teleosauroid relationships and diversity was better studied in the 21st century, the division of teleosauroids into two distinct evolutionary lineages led to the establishment of Teleosauridae as a more restrictive family within the group, together with its sister family
Machimosauridae Machimosauridae is an extinct family of teleosauroid thalattosuchian crocodyliforms. Text was copied from this source, which is available under Creative Commons Attribution 4.0 International License The family was first identified in 2016, wh ...
. Amongst teleosauroids, teleosaurids were generally smaller and less common than machimosaurids, suggesting the two families occupied different niches, similar to modern species of
crocodilians Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period (Cenomanian stage) and are the closest living re ...
. However, teleosaurids were more diverse than machimosaurids, with generalist coastal predators (''
Mystriosaurus ''Mystriosaurus'' is an extinct genus of teleosaurid crocodyliform from the Early Jurassic (Toarcian). Fossil specimens have been found in the Whitby Mudstone of England and Posidonia Shale of Germany. The only known species, ''M. laurillardi'', ...
''), long-snouted marine
piscivores A piscivore () is a carnivorous animal that eats primarily fish. The name ''piscivore'' is derived . Piscivore is equivalent to the Greek-derived word ichthyophage, both of which mean "fish eater". Fish were the diet of early tetrapod evolut ...
(''
Bathysuchus ''Bathysuchus'' ("deep water crocodile") is an extinct genus of teleosaurid thalattosuchian from Late Jurassic (Kimmeridgian) deep water marine deposits in England and France. ''Bathysuchus'' displays features that suggest it was more pelagic th ...
''), and potentially even long-snouted, semi-terrestrial predators (''
Teleosaurus ''Teleosaurus'' (from el, τέλειος , 'perfect' and el, σαῦρος , 'lizard') is an extinct genus of teleosaurid crocodyliform found in the Middle Jurassic Calcaire de Caen Formation of France. It was approximately in length and w ...
''). Additionally, teleosaurids occupied a wider range of habitats than machimosaurids, from semi-marine coasts and esturies, the open-ocean, freshwater, and potentially even semi-terrestrial environments.


Classification

Teleosauridae is
phylogenetically In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
defined as the largest
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
of teleosauroids containing ''Teleosaurus'' but not ''
Machimosaurus ''Machimosaurus'' is an extinct genus of machimosaurid crocodyliform from the Late Jurassic (Kimmeridgian and Tithonian) and Early Cretaceous. The type species, ''Machimosaurus hugii'', was found in Switzerland. Other fossils have been found ...
'' and '' Plagiophthalmosuchus''. Teleosauridae is split into two
subfamilies In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
, the Teleosaurinae and the Aeolodontinae.


Palaeobiology

Teleosaurids were originally regarded as marine analogues to modern
gharials Gavialidae is a family of large semiaquatic crocodilians with elongated, narrow snouts. Gavialidae consists of two living species, the gharial (''Gavialis gangeticus'') and the false gharial (''Tomistoma schlegelii''), both occurring in Asia. Man ...
, as they both typically share long, tubular snouts and narrow teeth. However, differences in the jaws, teeth, and skeleton of different teleosaurids suggest that they were more ecologically diverse than this. Earlier teleosaurids were coastal semi-aquatic generalists, while the two subfamilies were more specialised. Teleosaurines appear to have been semi-terrestrial, as they were more heavily armoured and had forward-facing nostrils. In contrast, aeolodontines have been found in deep marine waters and had reduced armour, implying that they were open water predators similar to metriorhynchoids (although the oldest aeolodontine, ''
Mycterosuchus ''Mycterosuchus'' is an extinct genus of teleosaurid crocodyliform from the Middle Jurassic (Callovian) of England. Although previously synonymized with ''Steneosaurus'', recent cladistic analysis considers it distantly related to the ''Sten ...
'', appears to have been semi-terrestrial, similar to teleosaurines).


Palaeoecology


Distribution

Definitive fossils of teleosaurids are restricted to Laurasia, with material found in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
(
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
,
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
,
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
,
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
and
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
) and Asia (
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
and
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
, and possibly
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
).


See also

*
List of marine reptiles Following is a list of marine reptiles, reptiles which are adapted to life in marine or brackish environments. Extant The following marine reptiles are species which are currently extant or recently extinct. Crocodiles :*'' Crocodylus'' ::' ...


References

Jurassic crocodylomorphs Prehistoric reptile families Prehistoric marine crocodylomorphs Toarcian first appearances Hauterivian extinctions {{paleo-archosaur-stub