Tawny-throated Leaftosser
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The tawny-throated leaftosser (''Sclerurus mexicanus'') is a
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
in the ovenbird
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Furnariidae. It is also known as the tawny-throated leafscraper, Middle American leaftosser, Mexican leaftosser or Mexican leafscraper.


Description

This roughly thrush-sized bird measures 15–17 cm in length and weighs 24–30 g. Its coloration is quite uniformly a rich chestnut brown. The tail is darker, the breast, rump and head are lighter and tinged rufous, though the crown and cheek region are as dark as the body, with some greyish hue to the cheeks. Its
iris Iris most often refers to: *Iris (anatomy), part of the eye *Iris (mythology), a Greek goddess * ''Iris'' (plant), a genus of flowering plants * Iris (color), an ambiguous color term Iris or IRIS may also refer to: Arts and media Fictional ent ...
is dusky brown, the feet are blackish brown. The bill is very long and thin, perhaps by proportion the longest and thinnest of all Furnariidae. It is blackish brown above, and whitish, horn-colored or dark grey below; the tip is black.Remsen (2003a) Males and females look alike. Young birds are duller, with light streaks and dusky scaling on throat and breast. The song of the tawny-throated leaftosser is a series of 4–9 wheezy notes which descend, accelerate and fade out as they are given: '. For some populations, slightly different songs have been described; subspecies ''pullus'' produces a series of sharp ' calls or a whistling ', while subspecies ''peruvianus'' has a series of clear ' whistles and its song often ends in a trill. It is not clear whether these vocalizations indicate specific distinctiveness, but it is notable that at least in ''peruvianus'' the song often ends with a "flourish" and does not quietly fade out as it does in the northern populations. The alarm call is a sharp ''chick'', ', ' or '. The
short-billed leaftosser The short-billed leaftosser (''Sclerurus rufigularis'') is a species of bird in the family Furnariidae. It is found in Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, and Venezuela. Its natural habitat is subtropical o ...
(''S. rufigularis'') occurs widely in the Amazonas basin. It looks almost alike but it has a shorter bill and its song, while structured similarly, changes in pitch several times through its course rather than simply descending. It is also an inhabitant of ''
terra firme Terra may often refer to: * Terra (mythology), primeval Roman goddess * An alternate name for planet Earth, as well as the Latin name for the planet Terra may also refer to: Geography Astronomy * Terra (satellite), a multi-national NASA scienti ...
'' forest near rivers, whereas the tawny-throated leaftosser prefers hilly terrain.Remsen (2003a,b)


Systematics

The tawny-throated leaftosser is considered to be the
sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...
of the
short-billed leaftosser The short-billed leaftosser (''Sclerurus rufigularis'') is a species of bird in the family Furnariidae. It is found in Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Suriname, and Venezuela. Its natural habitat is subtropical o ...
(''S. rufigularis'').


Subspecies

* ''Sclerurus mexicanus mexicanus'' P.L.Sclater, 1857
Veracruz Veracruz (), formally Veracruz de Ignacio de la Llave (), officially the Free and Sovereign State of Veracruz de Ignacio de la Llave ( es, Estado Libre y Soberano de Veracruz de Ignacio de la Llave), is one of the 31 states which, along with Me ...
(SE
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
) to N
Nicaragua Nicaragua (; ), officially the Republic of Nicaragua (), is the largest country in Central America, bordered by Honduras to the north, the Caribbean to the east, Costa Rica to the south, and the Pacific Ocean to the west. Managua is the cou ...
. First recorded in
El Salvador El Salvador (; , meaning " The Saviour"), officially the Republic of El Salvador ( es, República de El Salvador), is a country in Central America. It is bordered on the northeast by Honduras, on the northwest by Guatemala, and on the south b ...
on June 12, 1998, but nesting not confirmed. Includes ''S. m. certus''. :Neither dark nor rufous hues very pronounced, but dark and rufous areas clearly contrasting. * ''Sclerurus mexicanus pullus'' Bangs, 1902 – Highlands from
Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...
to Coclé and
Veraguas Veraguas () is a province of Panama, located in the centre-west of the country. The capital is the city of Santiago de Veraguas. It is the only Panamanian province to border both the Atlantic and Pacific oceans. It covers an area of 10,587.6&nb ...
(W
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...
). :Generally darker and less rufous than ''mexicanus'', but throat lighter and rump with pronounced rufous hue.


Range and ecology

The tawny-throated leaftosser ranges from southern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
through
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
into
Panama Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Cos ...
. The species is patchily distributed across most of its range. Its natural
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s are tropical moist lowland
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s and tropical moist montane evergreen forests. It prefers
hill A hill is a landform that extends above the surrounding terrain. It often has a distinct Summit (topography), summit. Terminology The distinction between a hill and a mountain is unclear and largely subjective, but a hill is universally con ...
y terrain, occurring mainly between 700 and 2,200 m
ASL American Sign Language (ASL) is a natural language that serves as the predominant sign language of Deaf communities in the United States of America and most of Anglophone Canada. ASL is a complete and organized visual language that is express ...
in Central America' The species lives on the ground, where it feeds on
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s found by flicking through
leaf litter Plant litter (also leaf litter, tree litter, soil litter, litterfall or duff) is dead plant material (such as leaves, bark, needles, twigs, and cladodes) that have fallen to the ground. This detritus or dead organic material and its constituent ...
, digging in moist soil or in rotting wood, often using the tail to anchor itself; the tips of the
rectrices Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tail ...
become abraded by this behavior and before
moult In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is the manner in which an animal routinely casts off a part of its body (often, but not always, an outer ...
often just the resilient feather shafts remain. Tawny-throated leaftossers are encountered singly or in pairs; they may be attracted by recordings of their alarm calls. These birds typically move by hopping about and are reluctant to fly if they do not have to. They are not migratory.Remsen (2003a), Cuervo ''et al.'' (2007) Tawny-throated leaftossers are assumed to be monogamous and are territorial, with their
territories A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or an ...
that are around 25 ha in size. The breeding season – if this species actually has a well-marked breeding season – is prolonged, running at least from December to April in
Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the no ...
. The nest is a small cup, loosely woven from dry twigs and leaf remains, in a cavity up to 20 cm in diameter at the end of a tunnel – up to 50 cm long but often much shorter – dug into an earth bank or similar spot. Little is known about their breeding biology, but they presumably lay two eggs per
clutch A clutch is a mechanical device that engages and disengages power transmission, especially from a drive shaft to a driven shaft. In the simplest application, clutches connect and disconnect two rotating shafts (drive shafts or line shafts). ...
.Remsen (2003a), Greeney ''et al.'' (2004) It is generally an uncommon bird, with a
population density Population density (in agriculture: standing stock or plant density) is a measurement of population per unit land area. It is mostly applied to humans, but sometimes to other living organisms too. It is a key geographical term.Matt RosenberPopul ...
usually less than (and often far less than) 5 individuals per square kilometer. It quickly disappears from forests that have been fragmented due to logging, and will not even tolerate
selective logging In forestry, fishing and mining, high grading refers to the selective harvesting of goods to keep only the most valuable items. The term is frequently associated with fraud, especially in mining. Forestry In forestry, high grading, also sometimes ...
well. Due to its large overall range, it is classified as a Species of
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
by the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
>


Footnotes


References

* * Cuervo, Andrés M.; Hernández-Jaramillo, Alejandro; Cortés-Herrera, José Oswaldo & Laverde, Oscar (2007): Nuevos registros de aves en la parte alta de la Serranía de las Quinchas, Magdalena medio, Colombia ew bird records from the highlands of Serranía de las Quinchas, middle Magdalena valley, Colombia ''Ornitología Colombiana'' 5: 94–98 panish with English abstractbr>PDF fulltext
* Freile, Juan F. & Chaves, Jaime A. (2004): Interesting distributional records and notes on the biology of bird species from a cloud forest reserve in north-west Ecuador. ''
Bulletin of the British Ornithologists' Club The ''Bulletin of the British Ornithologists' Club'' is an ornithological journal published by the British Ornithologists' Club (BOC). It is cited as ''Bull. B. O. C.'' Many descriptions of birds new to science have been published in the bulleti ...
'' 124(1): 6–16
PDF fulltext
* Greeney, Harold F.; Gelis, Rudolphe A. & White, Richard (2004): Notes on breeding birds from an Ecuadorian lowland forest. ''
Bulletin of the British Ornithologists' Club The ''Bulletin of the British Ornithologists' Club'' is an ornithological journal published by the British Ornithologists' Club (BOC). It is cited as ''Bull. B. O. C.'' Many descriptions of birds new to science have been published in the bulleti ...
'' 124(1): 28–37
PDF fulltext
* Herrera, Néstor; Rivera, Roberto; Ibarra Portillo, Ricardo & Rodríguez, Wilfredo (2006): Nuevos registros para la avifauna de El Salvador. New records for the avifauna of El Salvador" ''Boletín de la Sociedad Antioqueña de Ornitología'' 16(2): 1–19. [Spanish with English abstract
PDF fulltext
* Remsen, J. Van (2003a): 223. Tawny-throated Leaftosser. ''In:'' del Hoyo, Josep; Elliott, Andrew & Christie, David A. (eds.): ''
Handbook of Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. Th ...
'' (Vol. 8: Broadbills to Tapaculos): 351, plate 29. Lynx Edicions, Barcelona. * Remsen, J. Van (2003b): 224. Short-billed Leaftosser. ''In:'' del Hoyo, Josep; Elliott, Andrew & Christie, David A. (eds.): ''
Handbook of Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. Th ...
'' (Vol. 8: Broadbills to Tapaculos): 351, plate 29. Lynx Edicions, Barcelona. * Salaman, Paul G.W.; Stiles, F. Gary; Bohórquez, Clara Isabel; Álvarez-R., Mauricio; Umaña, Ana María; Donegan, Thomas M. & Cuervo, Andrés M. (2002): New and noteworthy bird records from the east slope of the andes of Colombia. ''Caldasia'' 24(1): 157–189
PDF fulltext
{{Taxonbar, from=Q782336 tawny-throated leaftosser Birds of Central America Birds of Mexico Birds of Costa Rica Birds of Panama tawny-throated leaftosser tawny-throated leaftosser Taxonomy articles created by Polbot