The golden tanager (''Tangara arthus'') is a species of

bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...

in the family

Thraupidae The tanagers (singular ) comprise the bird family Thraupidae, in the order Passeriformes. The family has a Neotropical distribution and is the second-largest family of birds. It represents about 4% of all avian species and 12% of the Neotropica ...

. It is widespread and often common in highland forests of the

Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...

(from

Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...

and northwards) and

Venezuelan Coastal Range The Venezuelan Coastal Range ( es, Cordillera de la Costa or ), also known as Venezuelan Caribbean Mountain System ( es, Sistema Montañoso Caribe) is a mountain range system and one of the eight natural regions of Venezuela, that runs along the c ...

in north-western

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...

. Its

plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...

is overall golden-yellow with black to the back, wings, tail and ear-coverts. Some

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

are partially/largely brown below.

Taxonomy and systematics

The golden tanager was first described as ''Tangara Arthus'' by

René Lesson René-Primevère Lesson (20 March 1794 – 28 April 1849) was a French surgeon, naturalist, ornithologist, and herpetologist. Biography Lesson was born at Rochefort, and entered the Naval Medical School in Rochefort at the age of sixteen. He ...

in 1840 on the basis of a specimen from

Caracas Caracas (, ), officially Santiago de León de Caracas, abbreviated as CCS, is the capital and largest city of Venezuela, and the center of the Metropolitan Region of Caracas (or Greater Caracas). Caracas is located along the Guaire River in the ...

Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...

. The generic name ''Tangara'' comes from the

Tupí Tupí, also known as ''formatge de tupí'', is a fermented cheese of a certain area of the Pyrenees and Pre-Pyrenees made from cows' or sheep's milk. It is a cheese traditionally prepared in the mountainous Pallars region, as well as in the Cerda ...

word ''tangara'', meaning dancer. The specific name ''arthus'' is in honor of Arthus Bertrand, a French bookseller. Golden tanager is the official

common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...

designated by the

International Ornithologists' Union The International Ornithologists' Union, formerly known as the International Ornithological Committee, is a group of about 200 international ornithologists, and is responsible for the International Ornithological Congress and other international ...

(IOC). Other names for the species include "chestnut-breasted tanager". The golden tanager is one of 27 species in the genus '' Tangara''. Within the genus, it is part of a species group with the

blue-whiskered tanager The blue-whiskered tanager (''Tangara johannae'') is a species of bird in the family Thraupidae. It is found in the Chocó of Colombia and Ecuador. Its natural habitat is subtropical or tropical moist lowland forests. It is threatened by habitat ...

, green-and-gold tanager, emerald tanager, silver-throated tanager, saffron-crowned tanager, golden-eared tanager, and

flame-faced tanager The flame-faced tanager (''Tangara parzudakii'') is a species of bird in the tanager family Thraupidae. It is endemic to South America and is found in the eastern Andes of Colombia, Ecuador, Peru and Venezuela. Its natural habitat is subtropical ...

. In the group, it is

sister A sister is a woman or a girl who shares one or more parents with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a familial relationship, it is sometimes used endearingly to refer to ...

to a

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

formed by the emerald and silver-throated tanagers. This placement is supported by

mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...

evidence. The following

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...

shows

phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...

relationships within the species group based on the above study:

Subspecies

There are nine recognized subspecies of the golden tanager. The subspecies are differentiated by differences in their appearance and their distribution. All the subspecies excluding ''arthus'' are sometimes separated into a distinct species, ''Tangara aurulenta'', on the basis of differences in plumage. * ''T. a. arthus'' (

Lesson A lesson or class is a structured period of time where learning is intended to occur. It involves one or more students (also called pupils or learners in some circumstances) being taught by a teacher or instructor. A lesson may be either one ...

, 1832): The

nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

. It is found in the mountains of northern and western

. * ''T. a. palmitae'' ( Meyer de Schauensee, 1947): It is found on the western slope of the

in La Palmita,

Santander Santander may refer to: Places * Santander, Spain, a port city and capital of the autonomous community of Cantabria, Spain * Santander Department, a department of Colombia * Santander State, former state of Colombia * Santander de Quilichao, a m ...

Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...

. * ''T. a. sclateri'' ( Lafresnaye, 1854): It is found in the Andes of eastern Colombia, and likely also in those of southern

Táchira Táchira State ( es, Estado Táchira, ) is one of the 24 states of Venezuela. The state capital is San Cristóbal. Táchira State covers a total surface area of and as of the 2011 census, had a population of 1,168,908. At the end of the 19t ...

, Venezuela. * ''T. a. aurulenta'' (Lafresnaye, 1843): It is found in

Serranía del Perijá The Serranía del Perijá, Cordillera de Perijá or Sierra de Perijá is a mountain range, an extension of the eastern Andean branch ( Cordillera Oriental), in northern South America, between Colombia and Venezuela Venezuela (; ), offici ...

, Venezuela, and the western slope of the Andes in eastern Colombia. * ''T. a. occidentalis''

Chapman Chapman may refer to: Businesses * Chapman Entertainment, a former British television production company * Chapman Guitars, a guitar company established in 2009 by Rob Chapman * Chapman's, a Canadian ice cream and ice water products manufacturer ...

, 1914: It is found on the western slope of the Andes in central Colombia and on both slopes in western Colombia. * ''T. a. goodsoni''

Hartert Ernst Johann Otto Hartert (29 October 1859 – 11 November 1933) was a widely published German ornithologist. Life and career Hartert was born in Hamburg, Germany on 29 October 1859. In July 1891, he married the illustrator Claudia Bernadine E ...

, 1913: It is found on the western slope of the Andes in

Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ''Eku ...

, and likely also in northwestern

Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...

. * ''T. a. aequatorialis'' (

Taczanowski Taczanowski (Polish feminine: Taczanowska; plural: Taczanowscy) is the surname of a Polish szlachta (nobility) family from Poznań bearing the Jastrzębiec coat of arms and the motto: ''Plus penser que dire''. They took their name from th ...

& Berlepsch, 1885): It is found on the eastern slope of the Andes in Ecuador and adjoining areas in northern Peru. * ''T. a. pulchra'' ( Tschudi, 1844): It is found on the eastern slope of the Andes from Amazonas to Junín in Peru. * ''T. a. sophiae'' (Berlepsch, 1901): It is found on the eastern slope of the Andes from southeastern Peru to

La Paz La Paz (), officially known as Nuestra Señora de La Paz (Spanish pronunciation: ), is the seat of government of the Bolivia, Plurinational State of Bolivia. With an estimated 816,044 residents as of 2020, La Paz is the List of Bolivian cities ...

and

Cochabamba Cochabamba ( ay, Quchapampa; qu, Quchapampa) is a city and municipality in central Bolivia in a valley in the Andes mountain range. It is the capital of the Cochabamba Department and the fourth largest city in Bolivia, with a population of 63 ...

Distribution and habitat

The golden tanager is found in the

in Venezuela and in the Andes in

, and Bolivia at elevations of , but is most common at elevations of . It inhabits humid

montane Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...

evergreen forest An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, Live oak, acacias, magnolia, and banksia in more temperate zone ...

, as well as forest edges and nearby

secondary growth In botany, secondary growth is the growth that results from cell division in the cambia or lateral meristems and that causes the stems and roots to thicken, while primary growth is growth that occurs as a result of cell division at the tips of ...

Status

The nominate subspecies of the golden tanager is listed as being a species of

least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...

by the

International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

(IUCN) on the

IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...

because of its large range, relative commonness, and lack of a sufficiently rapid decline in population. The other subspecies, which are considered to be a distinct species by the IUCN, are also listed as being of least concern for the above reasons. However, the population of the golden tanager is decreasing, and it is threatened by

habitat destruction Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...

References

golden tanager Birds of the Northern Andes golden tanager Taxa named by René Lesson Taxonomy articles created by Polbot {{Thraupidae-stub