''Tagiades japetus'', commonly known as the pied flat or the common snow flat, is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of spread-winged skipper

butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...

belonging to the family

Hesperiidae Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy ...

. It is widely distributed, being found from India, the

Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...

, Southeast Asia, to Australia. It contains several recognized

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

Description

Life cycle and ecology

The eggs are laid on the upper surface of young leaves. They hatch in about six days, whereupon they will construct a leaf shelter by cutting the edge of a leaf, folding a triangular piece back, and then attaching it with silk. During the day, they hide under this shelter and only emerge at night to feed. The larvae feed on leaves of vines belonging to the genus ''

Dioscorea ''Dioscorea'' is a genus of over 600 species of flowering plants in the family Dioscoreaceae, native throughout the tropical and warm temperate regions of the world. The vast majority of the species are tropical, with only a few species extending ...

''; including ''

Dioscorea transversa ''Dioscorea transversa'', the pencil yam, is a vine of eastern and northern Australia. The leaves are heart-shaped, shiny, with 5-7 prominent veins. The seed pods are rounded, green or pink before drying to a straw brown papery texture. The ed ...

'', ''

Dioscorea alata ''Dioscorea alata'', also known as purple yam, ube (, ), or greater yam, among many other names, is a species of yam (a tuber). The tubers are usually a vivid violet-purple to bright lavender in color (hence the common name), but some range in ...

'', and '' Dioscorea numularia''. They may make other shelters as they grow larger. After about 23 days, they pupate inside their final shelter, emerging as adults after 10 days. The adults are quick flyers, maintaining a height relatively close to ground, generally under large trees, often resting under the leaves. They are usually encountered resting underneath leaf surfaces. They feed on nectar from flowers during the morning. Mating is not seasonal and adults emerge all throughout the year. However, the population is greatest during the

wet season The wet season (sometimes called the Rainy season) is the time of year when most of a region's average annual rainfall occurs. It is the time of year where the majority of a country's or region's annual precipitation occurs. Generally, the sea ...

Distribution and habitat

''Tagiades japetus'' have a wide range of occurrence. They are found from Sri Lanka and India to the Himalayas, and

Indochina Mainland Southeast Asia, also known as the Indochinese Peninsula or Indochina, is the continental portion of Southeast Asia. It lies east of the Indian subcontinent and south of Mainland China and is bordered by the Indian Ocean to the west an ...

. They also occur throughout the

Maritime Southeast Asia Maritime Southeast Asia comprises the countries of Brunei, Indonesia, Malaysia, the Philippines, Singapore, and East Timor. Maritime Southeast Asia is sometimes also referred to as Island Southeast Asia, Insular Southeast Asia or Oceanic Sout ...

to the Philippines and down to Papua New Guinea and the surrounding islands, and northeastern Australia. They are commonly found in the edges of rainforests, vine thickets, and sometimes in cultivated lands.

Subspecies

''Tagiades japetus'' contains several subspecies including the ones listed below with their common names and areas of distribution: *''Tagiades japetus atticus'' (Fabricius, 1793) - common snow flat (

Madhya Pradesh Madhya Pradesh (, ; meaning 'central province') is a state in central India. Its capital is Bhopal, and the largest city is Indore, with Jabalpur, Ujjain, Gwalior, Sagar, and Rewa being the other major cities. Madhya Pradesh is the seco ...

Bengal Bengal ( ; bn, বাংলা/বঙ্গ, translit=Bānglā/Bôngô, ) is a geopolitical, cultural and historical region in South Asia, specifically in the eastern part of the Indian subcontinent at the apex of the Bay of Bengal, predom ...

Dehra Dun Dehradun () is the capital and the most populous city of the Indian state of Uttarakhand. It is the administrative headquarters of the eponymous district and is governed by the Dehradun Municipal Corporation, with the Uttarakhand Legislativ ...

Assam Assam (; ) is a state in northeastern India, south of the eastern Himalayas along the Brahmaputra and Barak River valleys. Assam covers an area of . The state is bordered by Bhutan and Arunachal Pradesh to the north; Nagaland and Manipur ...

, Myanmar) *''Tagiades japetus avienus'' Fruhstorfer, 1910 -

Key Island Key Island, with the adjacent Key Reef, is a granite island, with an area of 6 ha, in south-eastern Australia. It is part of Tasmania’s Long Island Group, lying in eastern Bass Strait west of Cape Barren Island in the Furneaux Group. The ketc ...

*''Tagiades japetus bandanus'' Fruhstorfer, 1910 -

Banda Island The Banda Islands ( id, Kepulauan Banda) are a volcanic group of ten small volcanic islands in the Banda Sea, about south of Seram Island and about east of Java, and constitute an administrative district (''kecamatan'') within the Central M ...

*''Tagiades japetus brasidas'' Doherty, 1891 -

Sumba Sumba ( id, Pulau Sumba) is an island in eastern Indonesia. It is one of the Lesser Sunda Islands and is in the province of East Nusa Tenggara. Sumba has an area of , and the population was 779,049 at the 2020 Census; the official estimate as a ...

, Indonesia *''Tagiades japetus engnanicus'' Fruhstorfer, 1910 -

Enggano Island Enggano Island is about 100 km (60 mi) southwest of Sumatra, Indonesia. It is one of the 92 officially listed outlying islands of Indonesia, though it can also be considered a barrier island of Sumatra. Enggano is about long from ea ...

, western

Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...

*''Tagiades japetus janetta'' Butler, 1870 - black and white flat (northern gulf and north-eastern coast of

Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...

Aru Islands The Aru Islands Regency ( id, Kabupaten Kepulauan Aru) is a group of about 95 low-lying islands in the Maluku Islands of eastern Indonesia. It also forms a regency of Maluku Province, with a land area of . At the 2011 Census the Regency had a po ...

Irian Jaya New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torres ...

, Papua New Guinea, and

Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capita ...

) *''Tagiades japetus japetus'' (Stoll, 1782) -

Ambon Ambon may refer to: Places * Ambon Island, an island in Indonesia ** Ambon, Maluku, a city on Ambon Island, the capital of Maluku province ** Governorate of Ambon, a colony of the Dutch East India Company from 1605 to 1796 * Ambon, Morbihan, a c ...

, Maluku, Indonesia *''Tagiades japetus navus'' (Fruhstorfer, 1910) - Sula and

Tukangbesi Islands Tukangbesi Islands, is a group of islands off the coast of Sulawesi immediately east of Buton island in the Banda Sea region, and part of Sulawesi Tenggara. "Tukang Besi" literally means "iron worker" or "blacksmith" in Indonesian. There is a T ...

*''Tagiades japetus obscurus'' Mabille, 1876 (Ceylon snow flat) - Sri Lanka,

Nilgiris The Nilgiri Mountains form part of the Western Ghats in northwestern Tamil Nadu, Southern Karnataka, and eastern Kerala in India. They are located at the trijunction of three states and connect the Western Ghats with the Eastern Ghats. At le ...

Shevaroys The Selvarayan hills, with the anglicised name Shevaroy Hills, are a towering mountain range (1620 m) near the city of Salem, in Tamil Nadu state, southern India. It is one of the major hill stations in Tamil Nadu and in the Eastern Ghats. The loc ...

, and the

Palni Hills The Palani Hills are a mountain range in the southern Indian states of Kerala and Tamil Nadu. The Palani Hills are an eastward extension of the Western Ghats ranges, which run parallel to the west coast of India. The Palani Hills adjoin the hi ...

of southern India *''Tagiades japetus prasnaja'' Fruhstorfer, 1910 -

Sulawesi Sulawesi (), also known as Celebes (), is an island in Indonesia. One of the four Greater Sunda Islands, and the world's eleventh-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Mindanao and the Sulu Ar ...

, Banggai, and

Selayar Island Selayar is the main island of the Selayar Islands (''Kepulauan Selayar''). It lies off the coast of Cape Bira of South Sulawesi Province. The Selayar Straits separate it from the mainland of Sulawesi. Its main city is Benteng, towns to the so ...

*''Tagiades japetus titus'' (Plötz, 1884) - Philippine common snow flat (endemic to the Philippines) *''Tagiades japetus xarea'' Mabille, 1891 -

Timor Timor is an island at the southern end of Maritime Southeast Asia, in the north of the Timor Sea. The island is East Timor–Indonesia border, divided between the sovereign states of East Timor on the eastern part and Indonesia on the western p ...

File:Tagiades japetus janetta.jpg, ''Tagiades japetus janetta'' File:Tagiades japetus (Mindanao, Philippines) 2.jpg, ''Tagiades japetus titus'' from the Philippines File:Tagiades japetus mating.JPG, ''Tagiades japetus obscurus'', mating

References

External links

Australian Faunal Directory
{{Taxonbar, from=Q7675268 Tagiades Butterflies of Singapore Butterflies of Indochina