Tachyeres
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The steamer ducks are a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
(''Tachyeres'') of
duck Duck is the common name for numerous species of waterfowl in the family Anatidae. Ducks are generally smaller and shorter-necked than swans and geese, which are members of the same family. Divided among several subfamilies, they are a form ...
s in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Anatidae The Anatidae are the biological family of water birds that includes ducks, geese, and swans. The family has a cosmopolitan distribution, occurring on all the world's continents except Antarctica. These birds are adapted for swimming, floating ...
. All of the four species occur at the southern cone of
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
in Chile and Argentina, and all except the flying steamer duck are flightless; even this one species capable of flight rarely takes to the air. They can be aggressive and are capable of chasing off predators like petrels. Bloody battles of steamer ducks with each other over territory disputes are observed in nature. They even kill waterbirds that are several times their size.


Taxonomy

The genus ''Tachyeres'' was introduced in 1875 by the English zoologist Richard Owen to accommodate the
Falkland steamer duck The Falkland steamer duck (''Tachyeres brachypterus'') is a species of flightless duck found on the Falkland Islands in the South Atlantic Ocean. The steamer ducks get their name from their unconventional swimming behaviour in which they flap the ...
. The genus name ''Tachyeres'', "having fast
oar An oar is an implement used for water-borne propulsion. Oars have a flat blade at one end. Rowers grasp the oar at the other end. The difference between oars and paddles is that oars are used exclusively for rowing. In rowing the oar is connecte ...
s" or "fast rower", comes from
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
ταχυ- "fast" + ἐρέσσω "I row (as with oars)". The common name "steamer ducks" arose because, when swimming fast, they flap their wings into the water as well as using their feet, creating an effect like a
paddle steamer A paddle steamer is a steamship or steamboat powered by a steam engine that drives paddle wheels to propel the craft through the water. In antiquity, paddle wheelers followed the development of poles, oars and sails, where the first uses we ...
. They are usually placed in the
shelduck The shelducks, most species of which are found in the genus ''Tadorna'' (except for the Radjah shelduck, which is now found in its own monotypic genus ''Radjah''), are a group of large birds in the Tadorninae subfamily of the Anatidae, the biolog ...
subfamily Tadorninae. However,
mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA ...
sequence In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is calle ...
analyses of the cytochrome ''b'' and
NADH dehydrogenase NADH dehydrogenase is an enzyme that converts nicotinamide adenine dinucleotide (NAD) from its reduced form (NADH) to its oxidized form (NAD+). Members of the NADH dehydrogenase family and analogues are commonly systematically named using the for ...
subunit 2 genes indicate that ''Tachyeres'' rather belongs in a distinct clade of aberrant South American dabbling ducks, which also includes the Brazilian, the crested, and the
bronze-winged duck The bronze-winged duck (''Speculanas specularis'') also known as the spectacled duck, is a dabbling duck and the sole member of its genus ''Speculanas''. It is often placed in ''Anas'' with most other dabbling ducks, but its closest relative is ...
s.


Extant species

There are four species: The Chubut steamer duck was only described in 1981.


Phylogeny

Based on the ''Taxonomy in Flux'' from John Boyd's website.


Evolution


Flightlessness

Flightless ''Tachyeres'' have a paraphyletic organization, as shown above. There are multiple possible explanations of these organizations. It is unlikely that flightlessness evolved once in all ''Tachyeres'' and then disappeared in ''T. patachonicus'', because there is no evidence for a reversal of evolution, and these reversals are extremely rare. It is more likely that flightlessness evolved independently in each steamer duck species. The
DYRK1A Dual specificity tyrosine-phosphorylation-regulated kinase 1A is an enzyme that in humans is encoded by the ''DYRK1A'' gene. Alternative splicing of this gene generates several transcript variants differing from each other either in the Five prime ...
enzyme has been identified as a candidate gene for flightlessness in steamer ducks. This finding, combined with the range of flight capability, means the evolutionary history of the group may not be so clear cut. There is genomic evidence of recent speciation into four ''Tachyeres'' species. Flightless ''Tachyeres'' are thought to be undergoing a modern evolutionary transition to flightlessness, which explains the range of flight capability observed across the genus. The largest males of the most volant species, the flying steamer duck, are completely incapable of flight, while other individuals rarely fly. The flying steamer duck is the only species to reside in landlocked bodies of water. Generally, island bound/isolated avian populations are more likely to experience evolution towards flightlessness, which may be the case for several ''Tachyeres'' populations in the coastal South American regions.


References


External links

* * Ducks * {{duck-stub