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''Sonorasaurus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomencla ...
of
brachiosaurid The Brachiosauridae ("arm lizards", from Greek ''brachion'' (βραχίων) = "arm" and ''sauros'' = "lizard") are a family or clade of herbivorous, quadrupedal sauropod dinosaurs. Brachiosaurids had long necks that enabled them to access the le ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23  million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is ...
from the Early to Late Cretaceous (
Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
to Cenomanian
stages Stage or stages may refer to: Acting * Stage (theatre), a space for the performance of theatrical productions * Theatre, a branch of the performing arts, often referred to as "the stage" * ''The Stage'', a weekly British theatre newspaper * St ...
, around 112 to 93 
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
). It was a
herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpar ...
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
whose fossils have been found in southern Arizona in the United States. Its name, which means "Sonora lizard", comes from the
Sonoran Desert The Sonoran Desert ( es, Desierto de Sonora) is a desert in North America and ecoregion that covers the northwestern Mexican states of Sonora, Baja California, and Baja California Sur, as well as part of the southwestern United States (in Arizona ...
where its fossils were first found. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
is ''S. thompsoni'', described by Ratkevich in 1998.


Discovery

Fossilized remains were discovered in November 1994 by geology student Richard Thompson in the Turney Ranch Formation, located in the
Chihuahua Desert Chihuahua may refer to: Places *Chihuahua (state), a Mexican state **Chihuahua (dog), a breed of dog named after the state ** Chihuahua cheese, a type of cheese originating in the state **Chihuahua City, the capital city of the state ** Chihuahua M ...
region of the
Sonoran Desert The Sonoran Desert ( es, Desierto de Sonora) is a desert in North America and ecoregion that covers the northwestern Mexican states of Sonora, Baja California, and Baja California Sur, as well as part of the southwestern United States (in Arizona ...
in southern Arizona. Thompson had investigated a previously almost unexplored region, where fossils proved to be plentiful and directly accessible on the surface. A relatively complete sauropod skeleton was weathering out on a rock wall. He informed paleontologist
Ronald Paul Ratkevich Ronald is a masculine given name derived from the Old Norse ''Rögnvaldr'', Hanks; Hardcastle; Hodges (2006) p. 234; Hanks; Hodges (2003) § Ronald. or possibly from Old English '' Regenweald''. In some cases ''Ronald'' is an Anglicised form of ...
of the
Arizona-Sonora Desert Museum The Arizona-Sonora Desert Museum is a 98-acre (40 ha) zoo, aquarium, botanical garden, natural history museum, publisher, and art gallery founded in 1952. Located just west of Tucson, Arizona, it features two miles (3.2 km) of walking paths ...
,
Tucson , "(at the) base of the black ill , nicknames = "The Old Pueblo", "Optics Valley", "America's biggest small town" , image_map = , mapsize = 260px , map_caption = Interactive map ...
of the find. Ratkevich assembled a team of volunteers and began securing the bones in the spring of 1995; excavations would only end in 1999. He assumed the fossils represented a sauropod but was no expert on that taxon. The curator of geology of the museum, David W. Thayer, thought it might be a
therizinosaur Therizinosaurs (once called segnosaurs) were large herbivorous theropod dinosaurs whose fossils have been found across the Early to Late Cretaceous deposits in Asia and North America. Various features of the forelimbs, skull and pelvis unite t ...
, mistaking a tail chevron bone for the long hand claw typical of that group. In 1995, Ratkevich and Tayer first reported the find, already using the name "Sonorasaurus" but informally, so that it remained a ''
nomen nudum In taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate descr ...
''. Both men now asked dinosaur expert
Edwin Harris Colbert Edwin Harris "Ned" Colbert (September 28, 1905 – November 15, 2001)O'Connor, Anahad ''The New York Times'', November 25, 2001. was a distinguished American vertebrate paleontologist and prolific researcher and author. Born in Clarinda, Iowa, he ...
to identify the animal. Colbert, having seen only pictures, suggested it might be a member of the
Hadrosauridae Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which includ ...
. Ratkevich and Thayer then visited the displays in the
American Museum of Natural History The American Museum of Natural History (abbreviated as AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. In Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 26 inte ...
, concluding their find was rather dissimilar to the hadrosaurid skeletons shown there, so that it must represent a species new to science. Ratkevich considered naming it "Chihuahuasaurus" but ultimately shied away from the comical contrast between the gigantic sauropod and the minute dog breed. In 1996, a subsequent article tried to fit the bones found, in a diagram of the hadrosaurid ''
Kritosaurus ''Kritosaurus'' is an incompletely known genus of hadrosaurid (duck-billed) dinosaur. It lived about 74.5-66 million years ago, in the Late Cretaceous of North America. The name means "separated lizard" (referring to the arrangement of the cheek ...
''. This attempt largely failed, with an ilium being mistaken for a shoulder blade. Again, the name "Sonorasaurus" was used but still invalidly. ''Sonorasaurus'' was finally formally described in 1998 by Ratkevich, who identified it as a brachiosaurid sauropod. Dating of the specimen found it to be the earliest known brachiosaurid to have lived in the 'middle' Cretaceous
Period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...
of
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the C ...
. On April 10, 2018, ''Sonorasaurus'' was declared the state dinosaur of Arizona.


Description

The holotype of ''Sonorasaurus'', ASDM 500, is an incomplete skeleton consisting of various postcranial elements, many of which are fragmentary. A complete dorsal rib from the same horizon, ASDM 807, may also be referrable to ''S. thompsoni''. ''Sonorasaurus'' is estimated to have been about and tall, about one third of the size of ''
Brachiosaurus ''Brachiosaurus'' () is a genus of sauropod dinosaur that lived in North America during the Late Jurassic, about 154to 150million years ago. It was first described by American paleontologist Elmer S. Riggs in 1903 from fossils found in th ...
''.


Classification

Ratkevich initially identified ''Sonorasaurus'' as a brachiosaurid. However phylogenetic studies in the following years failed to find a consensus, with some finding it to lie within Brachiosauridae and others outside of it. In no analysis was the recovered phylogenetic position of ''Sonorasaurus'' strongly supported until D'Emic ''et al.'' (2016), which found ''Sonorasaurus'' to fall confidently within the Brachiosauridae. However the authors noted that additional data was still required to firmly establish its lower-level affinities.


References


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{{Taxonbar, from1=Q19683306, from2=Q957146 Brachiosaurs Early Cretaceous dinosaurs of North America Late Cretaceous dinosaurs of North America Fossil taxa described in 1998 Paleontology in Arizona Symbols of Arizona