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The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the
Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. T ...
Period, at million years ago ( Mya), to the beginning of the
Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, w ...
Period, Mya. The Silurian is the shortest period of the
Paleozoic The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ...
Era An era is a span of time defined for the purposes of chronology or historiography, as in the regnal eras in the history of a given monarchy, a calendar era used for a given calendar, or the geological eras defined for the history of Earth. Comp ...
. As with other
geologic period The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochr ...
s, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major
Ordovician–Silurian extinction events The Late Ordovician mass extinction (LOME), sometimes known as the end-Ordovician mass extinction or the Ordovician-Silurian extinction, is the first of the "big five" major mass extinction events in Earth's history, occurring roughly 443 Mya. It ...
when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution:
vascular plants Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They ...
emerged from more primitive land plants,
dikarya Dikarya is a subkingdom of Fungi that includes the divisions Ascomycota and Basidiomycota, both of which in general produce dikaryons, may be filamentous or unicellular, but are always without flagella. The Dikarya are most of the so-called "hig ...
n fungi started expanding and diversifying along with
glomeromycota Glomeromycota (often referred to as glomeromycetes, as they include only one class, Glomeromycetes) are one of eight currently recognized divisions within the kingdom Fungi, with approximately 230 described species. Members of the Glomeromyco ...
n fungi, and three groups of arthropods (
myriapods Myriapods () are the members of subphylum Myriapoda, containing arthropods such as millipedes and centipedes. The group contains about 13,000 species, all of them terrestrial. The fossil record of myriapods reaches back into the late Silurian, ...
,
arachnids Arachnida () is a class of joint-legged invertebrate animals (arthropods), in the subphylum Chelicerata. Arachnida includes, among others, spiders, scorpions, ticks, mites, pseudoscorpions, harvestmen, camel spiders, whip spiders and vine ...
and hexapods) became fully terrestrialized. A significant evolutionary milestone during the Silurian was the diversification of jawed fish and
bony fish Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartil ...
.


History of study

The Silurian system was first identified by British geologist
Roderick Murchison Sir Roderick Impey Murchison, 1st Baronet, (19 February 1792 – 22 October 1871) was a Scottish geologist who served as director-general of the British Geological Survey from 1855 until his death in 1871. He is noted for investigating and ...
, who was examining fossil-bearing sedimentary rock
strata In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as ...
in south
Wales Wales ( cy, Cymru ) is a country that is part of the United Kingdom. It is bordered by England to the east, the Irish Sea to the north and west, the Celtic Sea to the south west and the Bristol Channel to the south. It had a population in ...
in the early 1830s. He named the sequences for a Celtic tribe of Wales, the
Silures The Silures ( , ) were a powerful and warlike tribe or tribal confederation of ancient Britain, occupying what is now south east Wales and perhaps some adjoining areas. They were bordered to the north by the Ordovices; to the east by the Dob ...
, inspired by his friend
Adam Sedgwick Adam Sedgwick (; 22 March 1785 – 27 January 1873) was a British geologist and Anglican priest, one of the founders of modern geology. He proposed the Cambrian and Devonian period of the geological timescale. Based on work which he did on ...
, who had named the period of his study the
Cambrian The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ag ...
, from the
Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
name for Wales. This naming does not indicate any correlation between the occurrence of the Silurian rocks and the land inhabited by the Silures ( . Geologic map of Wales, Map of pre-Roman tribes of Wales). In 1835 the two men presented a joint paper, under the title ''On the Silurian and Cambrian Systems, Exhibiting the Order in which the Older Sedimentary Strata Succeed each other in England and Wales,'' which was the germ of the modern
geological time scale The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochr ...
. As it was first identified, the "Silurian" series when traced farther afield quickly came to overlap Sedgwick's "Cambrian" sequence, however, provoking furious disagreements that ended the friendship.
Charles Lapworth Charles Lapworth FRS FGS (20 September 1842 – 13 March 1920) was a headteacher and an English geologist who pioneered faunal analysis using index fossils and identified the Ordovician period. Biography Charles Lapworth was born at Fari ...
resolved the conflict by defining a new
Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. T ...
system including the contested beds. An alternative name for the Silurian was ''"Gotlandian"'' after the strata of the Baltic island of Gotland. The French geologist Joachim Barrande, building on Murchison's work, used the term ''Silurian'' in a more comprehensive sense than was justified by subsequent knowledge. He divided the Silurian rocks of
Bohemia Bohemia ( ; cs, Čechy ; ; hsb, Čěska; szl, Czechy) is the westernmost and largest historical region of the Czech Republic. Bohemia can also refer to a wider area consisting of the historical Lands of the Bohemian Crown ruled by the Bohe ...
into eight stages. His interpretation was questioned in 1854 by
Edward Forbes Edward Forbes FRS, FGS (12 February 1815 – 18 November 1854) was a Manx naturalist. In 1846, he proposed that the distributions of montane plants and animals had been compressed downslope, and some oceanic islands connected to the mainlan ...
, and the later stages of Barrande; F, G and H have since been shown to be Devonian. Despite these modifications in the original groupings of the strata, it is recognized that Barrande established Bohemia as a classic ground for the study of the earliest Silurian fossils.


Subdivisions


Paleogeography

With the
supercontinent In geology, a supercontinent is the assembly of most or all of Earth's continent, continental blocks or cratons to form a single large landmass. However, some geologists use a different definition, "a grouping of formerly dispersed continents", ...
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period The Jurassic ( ) is a geologic period and st ...
covering the equator and much of the southern hemisphere, a large ocean occupied most of the northern half of the globe. The high sea levels of the Silurian and the relatively flat land (with few significant mountain belts) resulted in a number of island chains, and thus a rich diversity of environmental settings. During the Silurian, Gondwana continued a slow southward drift to high southern latitudes, but there is evidence that the Silurian icecaps were less extensive than those of the late-Ordovician glaciation. The southern continents remained united during this period. The melting of icecaps and
glacier A glacier (; ) is a persistent body of dense ice that is constantly moving under its own weight. A glacier forms where the accumulation of snow exceeds its ablation over many years, often centuries. It acquires distinguishing features, such as ...
s contributed to a rise in sea level, recognizable from the fact that Silurian sediments overlie eroded Ordovician sediments, forming an
unconformity An unconformity is a buried erosional or non-depositional surface separating two rock masses or strata of different ages, indicating that sediment deposition was not continuous. In general, the older layer was exposed to erosion for an interval ...
. The continents of
Avalonia Avalonia was a microcontinent in the Paleozoic era. Crustal fragments of this former microcontinent underlie south-west Great Britain, southern Ireland, and the eastern coast of North America. It is the source of many of the older rocks of Wester ...
,
Baltica Baltica is a paleocontinent that formed in the Paleoproterozoic and now constitutes northwestern Eurasia, or Europe north of the Trans-European Suture Zone and west of the Ural Mountains. The thick core of Baltica, the East European Craton, is ...
, and
Laurentia Laurentia or the North American Craton is a large continental craton that forms the ancient geological core of North America. Many times in its past, Laurentia has been a separate continent, as it is now in the form of North America, although ...
drifted together near the
equator The equator is a circle of latitude, about in circumference, that divides Earth into the Northern and Southern hemispheres. It is an imaginary line located at 0 degrees latitude, halfway between the North and South poles. The term can al ...
, starting the formation of a second supercontinent known as
Euramerica Laurasia () was the more northern of two large landmasses that formed part of the Pangaea supercontinent from around ( Mya), the other being Gondwana. It separated from Gondwana (beginning in the late Triassic period) during the breakup of Pa ...
. When the proto-Europe collided with North America, the collision folded coastal sediments that had been accumulating since the Cambrian off the east coast of North America and the west coast of Europe. This event is the
Caledonian orogeny The Caledonian orogeny was a mountain-building era recorded in the northern parts of the British Isles, the Scandinavian Mountains, Svalbard, eastern Greenland and parts of north-central Europe. The Caledonian orogeny encompasses events that ...
, a spate of mountain building that stretched from
New York State New York, officially the State of New York, is a state in the Northeastern United States. It is often called New York State to distinguish it from its largest city, New York City. With a total area of , New York is the 27th-largest U.S. sta ...
through conjoined Europe and Greenland to Norway. At the end of the Silurian, sea levels dropped again, leaving telltale basins of
evaporite An evaporite () is a water-soluble sedimentary mineral deposit that results from concentration and crystallization by evaporation from an aqueous solution. There are two types of evaporite deposits: marine, which can also be described as ocea ...
s extending from Michigan to West Virginia, and the new mountain ranges were rapidly eroded. The
Teays River The Teays River (pronounced taze) was a major preglacial river that drained much of the present Ohio River watershed, but took a more northerly downstream course. Traces of the Teays across northern Ohio and Indiana are represented by a network ...
, flowing into the shallow mid-continental sea, eroded Ordovician Period strata, forming deposits of Silurian strata in northern Ohio and Indiana. The vast ocean of
Panthalassa Panthalassa, also known as the Panthalassic Ocean or Panthalassan Ocean (from Greek "all" and "sea"), was the superocean that surrounded the supercontinent Pangaea, the latest in a series of supercontinents in the history of Earth. During ...
covered most of the northern hemisphere. Other minor oceans include two phases of the Tethys, the Proto-Tethys and Paleo-Tethys, the Rheic Ocean, the Iapetus Ocean (a narrow seaway between Avalonia and Laurentia), and the newly formed
Ural Ocean The Ural Ocean (also called the Uralic Ocean) was a small, ancient ocean that was situated between Siberia and Baltica. The ocean formed in the Late Ordovician epoch, when large islands from Siberia collided with Baltica, which was then part of ...
.


Climate and sea level

The Silurian period was once believed to have enjoyed relatively stable and warm temperatures, in contrast with the extreme glaciations of the Ordovician before it and the extreme heat of the ensuing Devonian; however, it is now known that the global climate underwent many drastic fluctuations throughout the Silurian, evidenced by numerous major carbon and oxygen isotope excursions during this geologic period. Sea levels rose from their
Hirnantian The Hirnantian is the final internationally recognized stage of the Ordovician Period of the Paleozoic Era. It was of short duration, lasting about 1.4 million years, from to Ma (million years ago). The early part of the Hirnantian was chara ...
low throughout the first half of the Silurian; they subsequently fell throughout the rest of the period, although smaller scale patterns are superimposed on this general trend; fifteen high-stands (periods when sea levels were above the edge of the continental shelf) can be identified, and the highest Silurian sea level was probably around higher than the lowest level reached. During this period, the
Earth Earth is the third planet from the Sun and the only astronomical object known to harbor life. While large volumes of water can be found throughout the Solar System, only Earth sustains liquid surface water. About 71% of Earth's surfa ...
entered a warm
greenhouse A greenhouse (also called a glasshouse, or, if with sufficient heating, a hothouse) is a structure with walls and roof made chiefly of transparent material, such as glass, in which plants requiring regulated climatic conditions are grown.These ...
phase, supported by high CO2 levels of 4500 ppm, and warm shallow seas covered much of the equatorial land masses. Early in the Silurian,
glacier A glacier (; ) is a persistent body of dense ice that is constantly moving under its own weight. A glacier forms where the accumulation of snow exceeds its ablation over many years, often centuries. It acquires distinguishing features, such as ...
s retreated back into the
South Pole The South Pole, also known as the Geographic South Pole, Terrestrial South Pole or 90th Parallel South, is one of the two points where Earth's axis of rotation intersects its surface. It is the southernmost point on Earth and lies antipod ...
until they almost disappeared in the middle of Silurian. Layers of broken shells (called
coquina Coquina () is a sedimentary rock that is composed either wholly or almost entirely of the transported, abraded, and mechanically sorted fragments of the shells of mollusks, trilobites, brachiopods, or other invertebrates. The term ''coqui ...
) provide strong evidence of a climate dominated by violent storms generated then as now by warm sea surfaces.


Perturbations

The climate and
carbon cycle The carbon cycle is the biogeochemical cycle by which carbon is exchanged among the biosphere, pedosphere, geosphere, hydrosphere, and atmosphere of the Earth. Carbon is the main component of biological compounds as well as a major component ...
appear to be rather unsettled during the Silurian, which had a higher frequency of isotopic excursions (indicative of climate fluctuations) than any other period. The Ireviken event, Mulde event and Lau event each represent isotopic excursions following a minor mass extinction and associated with rapid sea-level change. Each one leaves a similar signature in the geological record, both geochemically and biologically; pelagic (free-swimming) organisms were particularly hard hit, as were
brachiopods Brachiopods (), phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, ...
,
corals Corals are marine invertebrates within the class (biology), class Anthozoa of the phylum Cnidaria. They typically form compact Colony (biology), colonies of many identical individual polyp (zoology), polyps. Coral species include the important C ...
and
trilobites Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the ...
, and extinctions rarely occur in a rapid series of fast bursts. The climate fluctuations are best explained by a sequence of glaciations, but the lack of tillites in the middle to late Silurian make this explanation problematic.


Flora and fauna

The Silurian was the first period to see megafossils of extensive terrestrial biota in the form of
moss Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) ''sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and horn ...
-like miniature forests along lakes and streams and networks of large, mycorrhizal nematophytes, heralding the beginning of the Silurian-Devonian Terrestrial Revolution. However, the land fauna did not have a major impact on the Earth until it diversified in the Devonian. The first fossil records of
vascular plant Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They ...
s, that is, land plants with tissues that carry water and food, appeared in the second half of the Silurian Period. The earliest-known representatives of this group are ''
Cooksonia ''Cooksonia'' is an extinct group of primitive land plants, treated as a genus, although probably not monophyletic. The earliest ''Cooksonia'' date from the middle of the Silurian (the Wenlock epoch); the group continued to be an important c ...
''. Most of the sediments containing ''Cooksonia'' are marine in nature. Preferred habitats were likely along rivers and streams. ''
Baragwanathia ''Baragwanathia'' is a genus of extinct lycopsid plants of Late Silurian to Early Devonian age (), fossils of which have been found in Australia, Canada, China and Czechia. The name derives from William Baragwanath who discovered the first speci ...
'' appears to be almost as old, dating to the early Ludlow (420 million years) and has branching stems and needle-like leaves of . The plant shows a high degree of development in relation to the age of its fossil remains. Fossils of this plant have been recorded in Australia, Canada, and China. '' Eohostimella heathana'' is an early, probably terrestrial, "plant" known from compression fossils of Early Silurian (Llandovery) age. The chemistry of its fossils is similar to that of fossilised vascular plants, rather than algae. The earliest-known animals fully adapted to terrestrial conditions appear during the Mid-Silurian, including the millipede '' Pneumodesmus''. Some evidence also suggests the presence of predatory trigonotarbid arachnoids and
myriapod Myriapods () are the members of subphylum Myriapoda, containing arthropods such as millipedes and centipedes. The group contains about 13,000 species, all of them terrestrial. The fossil record of myriapods reaches back into the late Silurian, ...
s in Late Silurian facies. Predatory
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chorda ...
s would indicate that simple
food web A food web is the natural interconnection of food chains and a graphical representation of what-eats-what in an ecological community. Another name for food web is consumer-resource system. Ecologists can broadly lump all life forms into one ...
s were in place that included non-predatory prey animals. Extrapolating back from
Early Devonian The Early Devonian is the first of three epochs comprising the Devonian period, corresponding to the Lower Devonian series. It lasted from and began with the Lochkovian Stage , which was followed by the Pragian from and then by the Emsian, wh ...
biota, Andrew Jeram ''et al.'' in 1990 suggested a food web based on as-yet-undiscovered
detritivore Detritivores (also known as detrivores, detritophages, detritus feeders, or detritus eaters) are heterotrophs that obtain nutrients by consuming detritus (decomposing plant and animal parts as well as feces). There are many kinds of invertebrates, ...
s and grazers on micro-organisms. The first bony fish, the
Osteichthyes Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartilage ...
, appeared, represented by the Acanthodians covered with bony scales. Fish reached considerable diversity and developed movable
jaw The jaw is any opposable articulated structure at the entrance of the mouth, typically used for grasping and manipulating food. The term ''jaws'' is also broadly applied to the whole of the structures constituting the vault of the mouth and serv ...
s, adapted from the supports of the front two or three
gill A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they ar ...
arches. A diverse fauna of
eurypterid Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group i ...
s (sea scorpions)—some of them several meters in length—prowled the shallow Silurian seas of North America; many of their
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s have been found in
New York state New York, officially the State of New York, is a state in the Northeastern United States. It is often called New York State to distinguish it from its largest city, New York City. With a total area of , New York is the 27th-largest U.S. sta ...
.
Leech Leeches are segmented parasitic or predatory worms that comprise the subclass Hirudinea within the phylum Annelida. They are closely related to the oligochaetes, which include the earthworm, and like them have soft, muscular segmented bod ...
es also made their appearance during the Silurian Period. Brachiopods,
bryozoa Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about long, they have a special feeding structure called a ...
,
molluscs Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is es ...
, hederelloids, tentaculitoids,
crinoid Crinoids are marine animals that make up the class Crinoidea. Crinoids that are attached to the sea bottom by a stalk in their adult form are commonly called sea lilies, while the unstalked forms are called feather stars or comatulids, which ar ...
s and trilobites were abundant and diverse. Endobiotic symbionts were common in the corals and stromatoporoids. Reef abundance was patchy; sometimes, fossils are frequent, but at other points, are virtually absent from the rock record. File:Cooksonia.png, ''
Cooksonia ''Cooksonia'' is an extinct group of primitive land plants, treated as a genus, although probably not monophyletic. The earliest ''Cooksonia'' date from the middle of the Silurian (the Wenlock epoch); the group continued to be an important c ...
'', the earliest vascular plant, middle Silurian File:Wrens Nest Fossils 2.jpg, Silurian sea bed fossils collected from Wren's Nest Nature Reserve, Dudley UK File:Kaugatuma Bedding Plane Pridoli Estonia.jpg,
Crinoid Crinoids are marine animals that make up the class Crinoidea. Crinoids that are attached to the sea bottom by a stalk in their adult form are commonly called sea lilies, while the unstalked forms are called feather stars or comatulids, which ar ...
fragments in a Silurian (Pridoli) limestone (
Saaremaa Saaremaa is the largest island in Estonia, measuring . The main island of Saare County, it is located in the Baltic Sea, south of Hiiumaa island and west of Muhu island, and belongs to the West Estonian Archipelago. The capital of the island ...
, Estonia) File:Wrens Nest Fossils 3.jpg, Silurian sea bed fossils collected from Wren's Nest Nature Reserve, Dudley UK File:Eurypterus Paleoart.jpg, ''
Eurypterus ''Eurypterus'' ( ) is an extinct genus of eurypterid, a group of organisms commonly called "sea scorpions". The genus lived during the Silurian period, from around 432 to 418 million years ago. ''Eurypterus'' is by far the most well-studied and ...
'', a common Upper Silurian
eurypterid Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group i ...
File:Poraspis.jpg, '' Poraspis,'' a genus of armored
jawless fish Agnatha (, Ancient Greek 'without jaws') is an infraphylum of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both present ( cyclostomes) and extinct (conodonts and ostracoderms) species. Among recent animals, cyclostom ...
from the Late Silurian of
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world ...
,
Norway Norway, officially the Kingdom of Norway, is a Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of Jan Mayen and th ...
, and the
U.S The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territo ...
. File:Calymene celebra Raymond, 1916.jpg, Trilobites were still diverse and common in the Silurian. Fossils of ''Calymene celebra'' are extremely abundant in parts of central USA. File:HalysitesSilurian.jpg, ''
Halysites ''Halysites'' (meaning ''chain coral'') is an extinct genus of tabulate coral. Colonies range from less than one to tens of centimeters in diameter, and they fed upon plankton. These tabulate corals lived from the Ordovician to the Devonian (fro ...
'' was a
Tabulate coral Tabulata, commonly known as tabulate corals, are an order of extinct forms of coral. They are almost always colonial, forming colonies of individual hexagonal cells known as corallites defined by a skeleton of calcite, similar in appearance ...
, an extinct group that lived through the Paleozoic File:20211029 Parioscorpio venator.png, '' Parioscorpio'' was an enigmatic arthropod from the Silurian of
Wisconsin Wisconsin () is a state in the upper Midwestern United States. Wisconsin is the 25th-largest state by total area and the 20th-most populous. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake Michi ...
File:Dalmanites limulurus trilobite silurian.jpg, A ''
Dalmanites limulurus ''Dalmanites'' is a genus of trilobite in the order Phacopida. They lived from the Late Ordovician to Middle Devonian. The trilobites of this genus have slightly convex exoskeletons with an average length of . The cephalon is semicircular or ...
'' specimen from Silurian strata of
New York New York most commonly refers to: * New York City, the most populous city in the United States, located in the state of New York * New York (state), a state in the northeastern United States New York may also refer to: Film and television * ...
File:Geodized pentamerid brachiopods (Silurian; Swayzee, Indiana, USA) 1.jpg, A rock containing several geodized
pentamerid Pentamerida is an order of biconvex, impunctate shelled, articulate brachiopods that are found in marine sedimentary rocks that range from the Middle Cambrian through the Devonian.Moore, Lalcker and Fischer, 1952, Invertebrate Fossils, McGraw-Hil ...
brachiopods from strata in
Indiana Indiana () is a U.S. state in the Midwestern United States. It is the 38th-largest by area and the 17th-most populous of the 50 States. Its capital and largest city is Indianapolis. Indiana was admitted to the United States as the 19th st ...
File:Entelognathus.png, '' Entelognathus primordialis'' was a Placoderm fish from the late Silurian File:Jamoytius kerwoodi.jpg, '' Jamoytius'' was an enigmatic vertebrate that is possibly related to
Anaspid Anaspida ("without shield") is an extinct group of primitive jawless vertebrates that lived primarily during the Silurian period, and became extinct soon after the start of the Devonian. They were classically regarded as the ancestors of lampr ...
fish File:20201227 Pterygotus anglicus.png, ''
Pterygotus ''Pterygotus'' is a genus of giant predatory eurypterid, a group of extinct aquatic arthropods. Fossils of ''Pterygotus'' have been discovered in deposits ranging in age from Middle Silurian to Late Devonian, and have been referred to several di ...
'' was a giant eurypterid that had a nearly cosmopolitan distribution File:Sphooceras-truncatum.jpg, '' Sphooceras'' was a
Nautiloid Nautiloids are a group of marine cephalopods ( Mollusca) which originated in the Late Cambrian and are represented today by the living '' Nautilus'' and '' Allonautilus''. Fossil nautiloids are diverse and speciose, with over 2,500 recorded spec ...
cephalopod A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent hea ...
found in Silurian strata of the
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The Cz ...
File:Qianodus holotype.jpg, Qianodus is a tooth-based chondrichthyan genus from the early Silurian (
Aeronian In the geologic timescale, the Aeronian is an age of the Llandovery Epoch of the Silurian Period of the Paleozoic Era of the Phanerozoic Eon that began 440.8 ± 1.2 Ma and ended 438.5 ± 1.1 Ma (million years ago). The Aeronian Age succeed ...
) of China. File:Fanjingshania.jpg, Fanjingshania is a climatiid chondrichthyan from the lower Silurian (
Aeronian In the geologic timescale, the Aeronian is an age of the Llandovery Epoch of the Silurian Period of the Paleozoic Era of the Phanerozoic Eon that began 440.8 ± 1.2 Ma and ended 438.5 ± 1.1 Ma (million years ago). The Aeronian Age succeed ...
) described from disarticulated dermoskeletal elements.


Notes


References

* Emiliani, Cesare. (1992). ''Planet Earth : Cosmology, Geology, & the Evolution of Life & the Environment''. Cambridge University Press. (Paperback Edition ) * Mikulic, DG, DEG Briggs, and J Kluessendorf. 1985. A new exceptionally preserved biota from the Lower Silurian of Wisconsin, USA. Philosophical Transactions of the Royal Society of London, 311B:75-86. *


External links


Ogg, Jim (June 2004), ''Overview of Global Boundary Stratotype Sections and Points (GSSP's)''
Silurian

The Silurian
Paleoportal: Silurian strata in U.S., state by state
*

{{Authority control Geological periods