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Silage () is a type of fodder made from green foliage crops which have been preserved by fermentation to the point of acidification. It can be fed to
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Adult females are referred to as cows and adult ...
,
sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to domesticate ...
and other such ruminants (
cud Cud is a portion of food that returns from a ruminant's stomach to the mouth to be chewed for the second time. More precisely, it is a bolus of semi-degraded food regurgitated from the reticulorumen of a ruminant. Cud is produced during the p ...
-chewing animals). The fermentation and storage process is called ''ensilage'', ''ensiling'' or ''silaging''. Silage is usually made from
grass crops Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns ...
, including
maize Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American English, North American and Australian English), is a cereal grain first domesticated by indigenous peoples of Mexico, indigenous ...
, sorghum or other
cereal A cereal is any grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, germ, and bran. Cereal grain crops are grown in greater quantities and provide more food ...
s, using the entire green plant (not just the grain). Silage can be made from many field crops, and special terms may be used depending on type: ''oatlage'' for oats, ''haylage'' for alfalfa (''haylage'' may also refer to high dry matter silage made from hay). Silage can be made using several methods, largely dependent on available technology, local tradition or prevailing climate.


Production

The crops most often used for ensilage are the ordinary grasses,
clover Clover or trefoil are common names for plants of the genus ''Trifolium'' (from Latin ''tres'' 'three' + ''folium'' 'leaf'), consisting of about 300 species of flowering plants in the legume or pea family Fabaceae originating in Europe. The genus ...
s, alfalfa, vetches, oats, rye and
maize Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American English, North American and Australian English), is a cereal grain first domesticated by indigenous peoples of Mexico, indigenous ...
. Many crops have ensilaging potential, including
potato The potato is a starchy food, a tuber of the plant ''Solanum tuberosum'' and is a root vegetable native to the Americas. The plant is a perennial in the nightshade family Solanaceae. Wild potato species can be found from the southern Uni ...
es and various weeds, notably spurrey such as ''
Spergula arvensis ''Spergula arvensis'', the corn spurry, stickwort, starwort or spurrey, is a species of plant in the genus ''Spergula''. Description Corn spurry is a summer or winter annual broadleaf plant, and its seeds buried in the soil can survive for se ...
''. Silage must be made from plant material with a suitable moisture content: about 50% to 60% depending on the means of storage, the degree of compression, and the amount of water that will be lost in storage, but not exceeding 75%. Weather during harvest need not be as fair and dry as when harvesting for drying. For corn, harvest begins when the whole-plant moisture is at a suitable level, ideally a few days before it is ripe. For pasture-type crops, the grass is mown and allowed to wilt for a day or so until the moisture content drops to a suitable level. Ideally the crop is mowed when in full
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
, and deposited in the silo on the day of its cutting. After harvesting, crops are shredded to pieces about long. The material is spread in uniform layers over the floor of the silo, and closely packed. When the silo is filled or the stack built, a layer of
straw Straw is an agricultural byproduct consisting of the dry stalks of cereal plants after the grain and chaff have been removed. It makes up about half of the yield of cereal crops such as barley, oats, rice, rye and wheat. It has a number ...
or some other dry porous substance may be spread over the surface. In the silo, the pressure of the material, when
chaff Chaff (; ) is the dry, scaly protective casing of the seeds of cereal grains or similar fine, dry, scaly plant material (such as scaly parts of flowers or finely chopped straw). Chaff is indigestible by humans, but livestock can eat it. In agri ...
ed, excludes air from all but the top layer; in the case of the stack, extra
pressure Pressure (symbol: ''p'' or ''P'') is the force applied perpendicular to the surface of an object per unit area over which that force is distributed. Gauge pressure (also spelled ''gage'' pressure)The preferred spelling varies by country a ...
is applied by weights to prevent excessive heating.


Equipment

Forage harvesters collect and chop the plant material, and deposit it in trucks or wagons. These forage harvesters can be either
tractor A tractor is an engineering vehicle specifically designed to deliver a high tractive effort (or torque) at slow speeds, for the purposes of hauling a trailer or machinery such as that used in agriculture, mining or construction. Most commo ...
-drawn or self-propelled. Harvesters blow the chaff into the wagon through a chute at the rear or side of the machine. Chaff may also be emptied into a bagger, which puts the silage into a large plastic bag that is laid out on the ground. In North America, Australia, northwestern Europe, and New Zealand it is common for silage to be placed in large heaps on the ground, rolled by tractor to push out the air, then covered with plastic sheets that are held down by used tires or tire ring walls. In New Zealand and Northern Europe, 'clamps' made of concrete or old wooden railway ties (sleepers) and built into the side of a bank are sometimes used. The chopped grass can then be dumped in at the top, to be drawn from the bottom in winter. This requires considerable effort to compress the stack in the silo to cure it properly. Again, the pit is covered with plastic sheet and weighed down with tires. In an alternative method, the cut vegetation is formed into bales using a baler, making ''balage'' (North America) or ''silage bales'' (UK, Australia, New Zealand). The grass or other forage is cut and partly dried until it contains 30–40% moisture (much drier than bulk silage, but too damp to be stored as dry hay). It is then made into large bales which are wrapped tightly in plastic to exclude air. The plastic may wrap the whole of each cylindrical or cuboid bale, or be wrapped around only the curved sides of a cylindrical bale, leaving the ends uncovered. In this case, the bales are placed tightly end to end on the ground, making a long continuous "sausage" of silage, often at the side of a field. The wrapping may be performed by a
bale wrapper A bale wrapper is a farm implement for wrapping bales in plastic, for them to turn into silage. Bale wrappers come in three main forms - Turntable type, Satellite type and In-line Type. Satellite bale wrapper A satellite bale wrapper usually ...
, while the baled silage is handled using a bale handler or a front-loader, either impaling the bale on a flap, or by using a special grab. The flaps do not hole the bales. In the UK, baled silage is most often made in round bales about , individually wrapped with four to six layers of "bale wrap plastic" (black, white or green 25-micrometre stretch film). The percentage of dry matter can vary from about 20% dry matter upwards. The continuous "sausage" referred to above is made with a special machine which wraps the bales as they are pushed through a rotating hoop which applies the bale wrap to the outside of the bales (round or square) in a continuous wrap. The machine places the bales on the ground after wrapping by moving forward slowly during the wrapping process.


Haylage

''Haylage'' sometimes refers to high dry matter silage of around 40% to 60%, typically made from hay. Horse haylage is usually 60% to 70% dry matter, made in small bales or larger bales. Handling of wrapped bales is most often with some type of gripper that squeezes the plastic-covered bale between two metal parts to avoid puncturing the plastic. Simple fixed versions are available for round bales which are made of two shaped pipes or tubes spaced apart to slide under the sides of the bale, but when lifted will not let it slip through. Often used on the tractor rear three-point linkage, they incorporate a trip tipping mechanism which can flip the bales over on to the flat side or end for storage on the thickest plastic layers.


Fermentation

Silage undergoes anaerobic fermentation, which starts about 48 hours after the silo is filled, and converts sugars to acids. Fermentation is essentially complete after about two weeks. Before anaerobic fermentation starts, there is an aerobic phase in which the trapped
oxygen Oxygen is the chemical element with the symbol O and atomic number 8. It is a member of the chalcogen group in the periodic table, a highly reactive nonmetal, and an oxidizing agent that readily forms oxides with most elements ...
is consumed. How closely the fodder is packed determines the nature of the resulting silage by regulating the
chemical reaction A chemical reaction is a process that leads to the chemical transformation of one set of chemical substances to another. Classically, chemical reactions encompass changes that only involve the positions of electrons in the forming and breaking ...
s that occur in the stack. When closely packed, the supply of oxygen is limited, and the attendant acid fermentation brings about decomposition of the
carbohydrate In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may o ...
s present into acetic, butyric and lactic acids. This product is named sour silage. If the fodder is unchaffed and loosely packed, or the silo is built gradually,
oxidation Redox (reduction–oxidation, , ) is a type of chemical reaction in which the oxidation states of substrate change. Oxidation is the loss of electrons or an increase in the oxidation state, while reduction is the gain of electrons or ...
proceeds more rapidly and the temperature rises; if the mass is compressed when the temperature is , the action ceases and sweet silage results. The
nitrogen Nitrogen is the chemical element with the symbol N and atomic number 7. Nitrogen is a nonmetal and the lightest member of group 15 of the periodic table, often called the pnictogens. It is a common element in the universe, estimated at se ...
ous ingredients of the fodder also change: in making sour silage, as much as one-third of the
albuminoid Albumin is a family of globular proteins, the most common of which are the serum albumins. All the proteins of the albumin family are water-soluble, moderately soluble in concentrated salt solutions, and experience heat denaturation. Albumins ...
s may be converted into amino and ammonium compounds; in making sweet silage, a smaller proportion is changed, but they become less
digestible Digestion is the breakdown of large insoluble food molecules into small water-soluble food molecules so that they can be absorbed into the watery blood plasma. In certain organisms, these smaller substances are absorbed through the small intes ...
. If the fermentation process is poorly managed, sour silage acquires an unpleasant odour due to excess production of ammonia or butyric acid (the latter is responsible for the smell of rancid butter). In the past, the fermentation was conducted by indigenous microorganisms, but, today, some bulk silage is inoculated with specific microorganisms to speed fermentation or improve the resulting silage. Silage inoculants contain one or more strains of lactic acid bacteria, and the most common is '' Lactobacillus plantarum''. Other bacteria used include '' Lactobacillus buchneri'', ''
Enterococcus faecium ''Enterococcus faecium'' is a Gram-positive, gamma-hemolytic or non- hemolytic bacterium in the genus '' Enterococcus''. It can be commensal (innocuous, coexisting organism) in the gastrointestinal tract of humans and animals, but it may also ...
'' and '' Pediococcus'' species. Ryegrasses have high sugars and respond to nitrogen fertiliser better than any other grass species. These two qualities have made ryegrass the most popular grass for silage-making for the last sixty years. There are three ryegrasses in seed form and commonly used: Italian, Perennial and Hybrid.


Pollution and waste

The fermentation process of silo or pit silage releases liquid. Silo effluent is corrosive. It can also contaminate water sources unless collected and treated. The high nutrient content can lead to
eutrophication Eutrophication is the process by which an entire body of water, or parts of it, becomes progressively enriched with minerals and nutrients, particularly nitrogen and phosphorus. It has also been defined as "nutrient-induced increase in phyt ...
(hypertrophication), the growth of
bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were am ...
l or algal blooms. Plastic sheeting used for sealing pit or baled silage needs proper disposal, and some areas have recycling schemes for it. Traditionally, farms have burned silage plastics; however odor and smoke concerns have led certain communities to restrict that practice.


Storing silage

Silage must be firmly packed to minimize the oxygen content, lest it spoil. Silage goes through four major stages in a silo: *Presealing, which, after the first few days after filling a silo, enables some respiration and some dry matter (DM) loss, but stops. *Fermentation, which occurs over a few weeks. pH drops, and there is more DM loss, but hemicellulose is broken down; aerobic respiration stops. *Infiltration, which enables some oxygen infiltration, allowing for limited microbial respiration. Available carbohydrates (CHOs) are lost as heat and gas. *Emptying, which exposes surface, causing additional loss; rate of loss increases.


Safety

Silos are potentially hazardous: deaths may occur in the process of filling and maintaining them, and several safety precautions are necessary. There is a risk of injury by machinery or from falls. When a silo is filled, fine dust particles in the air can become explosive because of their large aggregate surface area. Also, fermentation presents respiratory hazards. The ensiling process produces "silo gas" during the early stages of the fermentation process. Silage gas contains
nitric oxide Nitric oxide (nitrogen oxide or nitrogen monoxide) is a colorless gas with the formula . It is one of the principal oxides of nitrogen. Nitric oxide is a free radical: it has an unpaired electron, which is sometimes denoted by a dot in its ...
(NO), which will react with oxygen (O2) in the air to form nitrogen dioxide (NO2), which is toxic. Lack of oxygen inside the silo can cause asphyxiation. Molds that grow when air reaches cured silage can cause
organic dust toxic syndrome Organic dust toxic syndrome is a potentially severe flu-like syndrome originally described in farmers, mushroom workers, bird breeders and other persons occupationally exposed to dusty conditions. Symptoms Symptoms arise 4 to 12 hours after expos ...
. Collapsing silage from large bunker silos has caused deaths. Silage itself poses no special danger.


Nutrition

Ensilage can be substituted for
root crop Root vegetables are underground plant parts eaten by humans as food. Although botany distinguishes true roots (such as taproots and tuberous roots) from non-roots (such as bulbs, corms, rhizomes, and tubers, although some contain both hypocotyl a ...
s. Bulk silage is commonly fed to dairy cattle, while baled silage tends to be used for beef cattle,
sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to domesticate ...
and
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
s. The advantages of silage as animal feed are several: *During fermentation, the silage bacteria act on the cellulose and carbohydrates in the forage to produce volatile fatty acids (VFAs), such as acetic, propionic, lactic, and butyric acids. By lowering pH, these produce a hostile environment for competing bacteria that might cause spoilage. The VFAs thus act as natural preservatives, in the same way that the lactic acid in yogurt and cheese increases the preservability of what began as milk, or vinegar (dilute acetic acid) preserves pickled vegetables. This preservative action is particularly important during winter in temperate regions, when green forage is unavailable. *When silage is prepared under optimal conditions, the modest acidity also has the effect of improving palatability and provides a dietary contrast for the animal. (However, excessive production of acetic and butyric acids can reduce palatability: the mix of bacteria is ideally chosen so as to maximize lactic acid production.) *Several of the fermenting organisms produce vitamins: for example, lactobacillus species produce
folic acid Folate, also known as vitamin B9 and folacin, is one of the B vitamins. Manufactured folic acid, which is converted into folate by the body, is used as a dietary supplement and in food fortification as it is more stable during processing a ...
and vitamin B12. *The fermentation process that produces VFA also yields energy that the bacteria use: some of the energy is released as heat. Silage is thus modestly lower in caloric content than the original forage, in the same way that yogurt has modestly fewer calories than milk. However, this loss of energy is offset by the preservation characteristics and improved digestibility of silage.


History

Using the same technique as the process for making
sauerkraut Sauerkraut (; , "sour cabbage") is finely cut raw cabbage that has been fermented by various lactic acid bacteria. It has a long shelf life and a distinctive sour flavor, both of which result from the lactic acid formed when the bacteria ...
, green fodder was preserved for animals in parts of Germany since the start of the 19th century. This gained the attention of French agriculturist Auguste Goffart of
Sologne Sologne (; ) is a natural region in Centre-Val de Loire, France, extending over portions of the departements of Loiret, Loir-et-Cher and Cher. Its area is about . To its north is the river Loire, to its south the river Cher, while the distri ...
, near
Orléans Orléans (;"Orleans"
(US) and
dairy farming in the United States suited the ensiling of green
corn Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. The ...
fodder, and was soon adopted by
New England New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
farmers. Francis Morris of Maryland prepared the first silage produced in America in 1876. The favourable results obtained in the US led to the introduction of the system in the United Kingdom, where Thomas Kirby first introduced the process for British dairy herds. The modern silage preserved with acid and by preventing contact with air was invented by Finnish academic and professor of chemistry Artturi Ilmari Virtanen. Virtanen was awarded the 1945 Nobel prize in chemistry "for his research and inventions in agricultural and nutrition chemistry, especially for his fodder preservation method", practically inventing modern silage. Early silos were made of stone or concrete either above or below ground, but it is recognized that air may be sufficiently excluded in a tightly pressed stack, though in this case a few inches of the fodder round the sides is generally useless owing to mildew. In the US, structures were typically constructed of wooden cylinders to 35 or 40 ft. in depth. In the early days of mechanized agriculture (late 1800s), stalks were cut and collected manually using a knife and horsedrawn wagon, and fed into a stationary machine called a "silo filler" that chopped the stalks and blew them up a narrow tube to the top of a tower silo.


Anaerobic digestion

Silage may be used for anaerobic digestion.


Fish silage

Fish silage is a method used for conserving
by-product A by-product or byproduct is a secondary product derived from a production process, manufacturing process or chemical reaction; it is not the primary product or service being produced. A by-product can be useful and marketable or it can be consid ...
s from fishing for later use as feed in
fish farming upright=1.3, Salmon farming in the sea (mariculture) at Loch Ainort, Isle of Skye">mariculture.html" ;"title="Salmon farming in the sea (mariculture">Salmon farming in the sea (mariculture) at Loch Ainort, Isle of Skye, Scotland Fish farming o ...
. This way, the parts of the fish that are not used as
human food Humans eat various food substances for enjoyment and nutritional support. It is usually of plant, animal, or fungal origin, and contains essential nutrients, such as carbohydrates, fats, proteins, vitamins, or minerals. Omnivorous humans are high ...
such as fish guts (entrails), fish heads and trimmings are utilized as ingredients in feed pellets. The silage is performed by first grinding the remains and mixing it with formic acid, and then storing it in a tank. The acid helps with
preservation Preservation may refer to: Heritage and conservation * Preservation (library and archival science), activities aimed at prolonging the life of a record while making as few changes as possible * ''Preservation'' (magazine), published by the Nat ...
as well as further dissolving the residues. Process tanks for fish silage can be aboard ships or on land.


See also

* Artturi Ilmari Virtanen * Grain crimping * Mycoestrogen *'' Trifolium pratense'' * Xenoestrogen


References


Sources

*''Making and Feeding Silage'', John Murdoch, B.Sc, Ph.D. Published by Dairy Farmer (Books) Limited, Lloyd's Chambers, Ipswich, UK 1961)
Feeding baleage to horses – the ultimate guide – Horsetalk.co.nz
* "The Owner-Built Homestead" by Barbara and Ken Kern, New York: Scribner, 1977.


Further reading

* Zhou, Yiqin. ''Compar son ofFresh or Ensiled Fodders (e.g., Grass, Legume, Corn) on the Production of Greenhouse Gases Following Enteric Fermentation in Beef Cattle''. Rouyn-Noranda, Qué.: Université du Québec en Abitibi-Témiscamingue, 2011. ''N.B''.: Research report. {{Authority control Anaerobic digestion Fodder