Siberian Stonechat
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The Siberian stonechat or Asian stonechat (''Saxicola maurus'') is a recently validated
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of the
Old World flycatcher The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, Bluethroat (''Luscinia svecica)'' and Norther ...
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
(Muscicapidae). Like the other thrush-like flycatchers, it was often placed in the
Turdidae The thrushes are a passerine bird family, Turdidae, with a worldwide distribution. The family was once much larger before biologists reclassified the former subfamily Saxicolinae, which includes the chats and European robins, as Old World flycat ...
in the past. It breeds in the East
Palearctic The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
including in easternmost Europe and winters in the
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by the ...
tropics The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
.


Description, systematics and taxonomy

It resembles its closest living relative the
European stonechat The European stonechat (''Saxicola rubicola'') is a small passerine bird that was formerly classed as a subspecies of the common stonechat. Long considered a member of the thrush family, Turdidae, genetic evidence has placed it and its relativ ...
(''S. rubicola''), but is typically darker above and paler below, with a white rump and whiter underparts with less orange on the breast. The male in breeding plumage has black upperparts and head (lacking the brownish tones of the European stonechat), a conspicuous white collar, scapular patch and rump, and a restricted area of orange on the throat.Robertson (1977), Stoddart (1992), Urquhart & Bowley (2002) The female has pale brown upperparts and head, white neck patches (not a full collar), and a pale, unstreaked pinkish-yellow rump. Males in winter plumage are intermediate between summer males and females, with a
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also ...
resembling the
whinchat The whinchat (''Saxicola rubetra'') is a small migratory passerine bird breeding in Europe and western Asia and wintering in central Africa. At one time considered to be in the thrush family, Turdidae, it is now placed in the Old World flycat ...
(''S. rubetra''); from this species and the female it can be distinguished by the full white collar. If seen at close distance, it can be recognized that its primary
remiges Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tail ...
are distinctly longer than in ''S. rubicola''. In this, it closely resembles the whinchat, which like ''S. maurus'' is adapted to long-distance
migration Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum le ...
s. The male has a clicking call, like two
pebble A pebble is a clast of rock with a particle size of based on the Udden-Wentworth scale of sedimentology. Pebbles are generally considered larger than granules ( in diameter) and smaller than cobbles ( in diameter). A rock made predominant ...
s knocked together. The song is high and twittering like the
dunnock The dunnock (''Prunella modularis'') is a small passerine, or perching bird, found throughout temperate Europe and into Asian Russia. Dunnocks have also been successfully introduced into New Zealand. It is by far the most widespread member of th ...
(''Prunella modularis''), an unrelated
passerida Passerida is, under the Sibley-Ahlquist taxonomy, one of two parvorders contained within the suborder Passeri (standard taxonomic practice would place them at the rank of infraorder). While more recent research suggests that its sister parvorder ...
n
songbird A songbird is a bird belonging to the suborder Passeri of the perching birds (Passeriformes). Another name that is sometimes seen as the scientific or vernacular name is Oscines, from Latin ''oscen'', "songbird". The Passeriformes contains 500 ...
belonging to the
Passeroidea Passerida is, under the Sibley-Ahlquist taxonomy, one of two parvorders contained within the suborder Passeri (standard taxonomic practice would place them at the rank of infraorder). While more recent research suggests that its sister parvorde ...
.Urquhart & Bowley (2002) There are five or six
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
, with ''S. m. maurus'' (described above) and the distinct but similar ''S. m. stejnegeri'' found across northern and central Asia. The southern ''S. m. variegatus'' (west of the
Caspian Sea The Caspian Sea is the world's largest inland body of water, often described as the world's largest lake or a full-fledged sea. An endorheic basin, it lies between Europe and Asia; east of the Caucasus, west of the broad steppe of Central Asia ...
), ''S. m. armenicus'' (eastern Turkey to Iran), ''S. m. indicus'' (
Himalaya The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
) and the Turkestan stonechat ''S. m. przewalskii'' (southwest China) are distinguished by larger white areas on the plumage. In the past, ''S. maurus'' was usually included in ''S. torquatus'' as part of the "
common stonechat Common stonechat is the name used for the ''Saxicola'' species ''Saxicola torquatus'' when this is treated in its broad sense. It is, however, now more widely considered to be a superspecies consisting of several related but distinct species, w ...
", but that
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
nowadays is restricted to the
African stonechat The African stonechat or common stonechat (''Saxicola torquatus'') is a species of the Old World flycatcher family (Muscicapidae), inhabiting sub-Saharan Africa and adjacent regions. Like the other chats, it was long assigned to the thrush fa ...
. Analysis of
mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA ...
cytochrome ''b''
sequence In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is calle ...
and
nDNA Nuclear DNA (nDNA), or nuclear deoxyribonucleic acid, is the DNA contained within each cell nucleus of a eukaryotic organism. It encodes for the majority of the genome in eukaryotes, with mitochondrial DNA and plastid DNA coding for the rest. I ...
microsatellite A microsatellite is a tract of repetitive DNA in which certain DNA motifs (ranging in length from one to six or more base pairs) are repeated, typically 5–50 times. Microsatellites occur at thousands of locations within an organism's genome. ...
fingerprinting A fingerprint is an impression left by the friction ridges of a human finger. The recovery of partial fingerprints from a crime scene is an important method of forensic science. Moisture and grease on a finger result in fingerprints on surfac ...
data, though not unequivocal, together with the evidence from
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
,
behaviour Behavior (American English) or behaviour (British English) is the range of actions and mannerisms made by individuals, organisms, systems or artificial entities in some environment. These systems can include other systems or organisms as wel ...
and
biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
seems to indicate that the present bird can be considered a distinct
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
. The European stonechat is its western
sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...
in the Eurasian lineage of stonechats; their ancestors separated during the
Late Pliocene Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, ...
or
Early Pleistocene The Early Pleistocene is an unofficial sub-epoch in the international geologic timescale in chronostratigraphy, being the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently estimated to span the time ...
, roughly 1.5-2.5
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
at the onset of the
Quaternary glaciation The Quaternary glaciation, also known as the Pleistocene glaciation, is an alternating series of glacial and interglacial periods during the Quaternary period that began 2.58 Ma (million years ago) and is ongoing. Although geologists describe ...
. Its scientific name means "dark rock-dweller". ''Saxicola'' derives from
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''saxum'' "rock" + ''incola'' "inhabitant"; ''maurus'' is Latinized from
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
''maúros'' (μαύρος) "black" (cf. "
moor Moor or Moors may refer to: Nature and ecology * Moorland, a habitat characterized by low-growing vegetation and acidic soils. Ethnic and religious groups * Moors, Muslim inhabitants of the Maghreb, Iberian Peninsula, Sicily, and Malta during ...
"), in reference to the upperpart colour as compared to ''S. rubicola''.


Distribution and ecology

The breeding range covers most of
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
, from about
latitude In geography, latitude is a coordinate that specifies the north– south position of a point on the surface of the Earth or another celestial body. Latitude is given as an angle that ranges from –90° at the south pole to 90° at the north pol ...
71°N in
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
south to the
Himalaya The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
and southwest China, and west to eastern Turkey and the
Caspian Sea The Caspian Sea is the world's largest inland body of water, often described as the world's largest lake or a full-fledged sea. An endorheic basin, it lies between Europe and Asia; east of the Caucasus, west of the broad steppe of Central Asia ...
area. It also breeds in the far northeast of Europe, mainly in Russia but occasionally as far west as Finland. The wintering range of the
migratory bird Bird migration is the regular seasonal movement, often north and south along a flyway, between Breeding in the wild, breeding and wintering grounds. Many species of bird migrate. Animal migration, Migration carries high costs in predation a ...
is from southern Japan south to Thailand and India, and west to northeast Africa. On migration, small numbers reach as far west as western Europe, and exceptionally as far east as Alaska in North America. The Siberian stonechat is insectivorous. It breeds in open rough scrubland or rough grassland with scattered
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s, from sea level to about 4,000 m
ASL American Sign Language (ASL) is a natural language that serves as the predominant sign language of Deaf communities in the United States of America and most of Anglophone Canada. ASL is a complete and organized visual language that is express ...
or more. The birds seem to avoid even cool
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
conditions and stay up north only during the hot continental summer. In the
montane Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
regions of the
Himalaya The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
foothills of Bhutan, migrants can on occasion be seen foraging in fields and pastures more than 2,000 m
ASL American Sign Language (ASL) is a natural language that serves as the predominant sign language of Deaf communities in the United States of America and most of Anglophone Canada. ASL is a complete and organized visual language that is express ...
, but most move further down and south to winter in
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
regions. Though it is not considered a distinct
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
by the
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
, it is widespread and common and would not be considered a threatened species.BLI (2009)


Gallery

Siberian stonechat (Saxicola maurus) male non-breeding.jpg, Male non-breeding,
Uttar Pradesh Uttar Pradesh (; , 'Northern Province') is a state in northern India. With over 200 million inhabitants, it is the most populated state in India as well as the most populous country subdivision in the world. It was established in 1950 ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
Asian stonechat (Saxicola maurus indicus) male non breeding Bardiya.jpg, Male ''S. m. indicus'',
Bardiya National Park ; , iucn_category = II , photo = Bardiya_02.jpg , photo_caption = , photo_alt= , map_image = , map_caption = Location in Nepal , location = Nepal , map = Nepal , relief = 1 , coordinates = , area_km2 = 968 , established = 1988 , go ...
,
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is mai ...
Saxicola stejnegeri, Hanoi, Vietnam 4G4A3231 (cropped).jpg, Siberian Stonechat Siberian Stonechat in Assam.jpg, Siberian Stonechat wintering at Dipor Bil, Assam, India


Footnotes


References

* Bangs, Outram (1932): Birds of western China obtained by the Kelley-Roosevelts expedition. '' Field Mus. Nat. Hist. Zool. Ser.'' 18(11): 343–379
Fulltext
at the
Internet Archive The Internet Archive is an American digital library with the stated mission of "universal access to all knowledge". It provides free public access to collections of digitized materials, including websites, software applications/games, music, ...
* * Inskipp, Carol; Inskipp, Tim & Sherub (2000)
The ornithological importance of Thrumshingla National Park, Bhutan
. ''
Forktail The forktails are small insectivorous birds in the genus ''Enicurus''. They were formerly in the thrush family, Turdidae, but are more often now treated as part of the Old World flycatcher family, Muscicapidae. Their name derives from their lo ...
'' 14: 147–162. * Robertson, Iain (1977): Identification and European status of eastern Stonechats '' Brit. Birds'' 70: 237–245. * Stoddart, Andy (1992): Identification of Siberian Stonechat ''
Birding World ''Birding World'' was a monthly birding magazine published in the United Kingdom. It was the magazine of the Bird Information Service, based at Cley next the Sea, Norfolk. With the publication of issue No. 26/12 in January 2014, ''Birding World'' ...
'' 5(9): 348–356. * Urquhart, Ewan & Bowley, Adam (2002): ''Stonechats. A Guide to the Genus Saxicola''.
Christopher Helm Christopher Alexander Roger Helm (born Dundee, 1 February 1937 – 20 January 2007) was a Scottish book publisher, notably of ornithology related titles, including the ''Helm Identification Guides''. Born in Dundee, he was raised in Forfar, w ...
, London. * Wink, M.; Sauer-Gürth, H. & Gwinner, E. (2002): Evolutionary relationships of stonechats and related species inferred from mitochondrial-DNA sequences and genomic fingerprinting. '' British Birds'' 95: 349–355
PDF fulltext
* Wittmann, U.; Heidrich, P.; Wink, M. & Gwinner, E. (1995): Speciation in the Stonechat (''Saxicola torquata'') inferred from nucleotide sequences of the cytochrome b-gene. ''Journal of Zoological Systematics and Evolutionary Research'' 33(2): 116–122. HTML abstract


External links


Xeno-canto: audio recordings of the Siberian stonechat
{{Taxonbar, from=Q1589281
Siberian stonechat The Siberian stonechat or Asian stonechat (''Saxicola maurus'') is a recently validated species of the Old World flycatcher family (Muscicapidae). Like the other thrush-like flycatchers, it was often placed in the Turdidae in the past. It breed ...
Siberian stonechat The Siberian stonechat or Asian stonechat (''Saxicola maurus'') is a recently validated species of the Old World flycatcher family (Muscicapidae). Like the other thrush-like flycatchers, it was often placed in the Turdidae in the past. It breed ...
Birds of North Asia
Siberian stonechat The Siberian stonechat or Asian stonechat (''Saxicola maurus'') is a recently validated species of the Old World flycatcher family (Muscicapidae). Like the other thrush-like flycatchers, it was often placed in the Turdidae in the past. It breed ...
Taxobox binomials not recognized by IUCN