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In
botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek w ...
, a plant shoot consists of any
plant stem A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, stores nutrien ...
together with its appendages, leaves and lateral buds, flowering stems, and
flower bud In botany, a bud is an undeveloped or embryonic shoot and normally occurs in the axil of a leaf or at the tip of a stem. Once formed, a bud may remain for some time in a dormant condition, or it may form a shoot immediately. Buds may be speci ...
s. The new growth from
seed germination Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, fern ...
that grows upward is a shoot where leaves will develop. In the spring,
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
plant shoots are the new growth that grows from the ground in
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
plants or the new stem or flower growth that grows on woody plants. In everyday speech, shoots are often synonymous with stems. Stems, which are an integral component of shoots, provide an axis for buds, fruits, and leaves. Young shoots are often eaten by animals because the fibers in the new growth have not yet completed secondary cell wall development, making the young shoots softer and easier to chew and digest. As shoots grow and age, the cells develop secondary cell walls that have a hard and tough structure. Some plants (e.g.
bracken Bracken (''Pteridium'') is a genus of large, coarse ferns in the family Dennstaedtiaceae. Ferns (Pteridophyta) are vascular plants that have alternating generations, large plants that produce spores and small plants that produce sex cells (eggs ...
) produce
toxin A toxin is a naturally occurring organic poison produced by metabolic activities of living cells or organisms. Toxins occur especially as a protein or conjugated protein. The term toxin was first used by organic chemist Ludwig Brieger (1849– ...
s that make their shoots inedible or less palatable. File:Cucumber leaf.jpg, The shoot of a
cucumber Cucumber (''Cucumis sativus'') is a widely-cultivated Vine#Horticultural climbing plants, creeping vine plant in the Cucurbitaceae family that bears usually cylindrical Fruit, fruits, which are used as culinary vegetables.
File:GiantKnotweed048.jpg, Edible shoots of Sachaline File:Sunflower seedlings.jpg,
Sunflower The common sunflower (''Helianthus annuus'') is a large annual forb of the genus ''Helianthus'' grown as a crop for its edible oily seeds. Apart from cooking oil production, it is also used as livestock forage (as a meal or a silage plant), as ...
seedlings
germinate Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, fer ...
File:Persea americana (Avocado) Sprout 08May2010.JPG, A young
hass avocado The Hass avocado is a variety of avocado with dark green, bumpy skin. It was first grown and sold by Southern California mail carrier and amateur horticulturist Rudolph Hass, who also gave it his name. The Hass avocado is a large-sized fruit weig ...
shoot


Shoot types of woody plants

Many
woody plant A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposite to herbaceous plants that die back to the ground until sp ...
s have distinct short shoots and long shoots. In some
angiosperm Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s, the short shoots, also called spur shoots or fruit spurs, produce the majority of flowers and fruit. A similar pattern occurs in some
conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
s and in ''
Ginkgo ''Ginkgo'' is a genus of non-flowering seed plants. The scientific name is also used as the English name. The order to which it belongs, Ginkgoales, first appeared in the Permian, 270 million years ago, and is now the only living genus with ...
'', although the "short shoots" of some genera such as ''Picea'' are so small that they can be mistaken for part of the leaf that they have produced. A related phenomenon is seasonal heterophylly, which involves visibly different leaves from spring growth and later
lammas growth Lammas growth, also called Lammas leaves, Lammas flush, second shoots, or summer shoots, is a season of renewed growth in some trees in temperate regions put on in July and August (if in the northern hemisphere, January and February if in the sout ...
. Whereas spring growth mostly comes from buds formed the previous season, and often includes flowers, lammas growth often involves long shoots. File: Suckers on stump.jpg, Long shoot growth File: Pyrus pyrifolia (Hosui) blossom.jpg, A mature fruiting spur on a Nashi pear tree, ''
Pyrus pyrifolia ''Pyrus pyrifolia'' is a species of pear tree native to East Asia. The tree's edible fruit is known by many names, including: Asian pear, Japanese pear, Chinese pear, Korean pear, Taiwanese pear, apple pear, zodiac pear, three-halves pear, pa ...
'' File: Cedrus deodara 02.JPG, On long shoots of ''
Cedrus deodara ''Cedrus deodara'', the deodar cedar, Himalayan cedar, or deodar, is a species of cedar native to the Himalayas. Description It is a large evergreen coniferous tree reaching tall, exceptionally with a trunk up to in diameter. It has a conic ...
'' individual leaves may have
bud In botany, a bud is an undeveloped or embryonic shoot and normally occurs in the axil of a leaf or at the tip of a stem. Once formed, a bud may remain for some time in a dormant condition, or it may form a shoot immediately. Buds may be spec ...
s in the axils. File:20130903Cedrus deodara2.jpg, ''Cedrus deodara'' forms short shoots (from buds) along the long shoots.


See also

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Bud In botany, a bud is an undeveloped or embryonic shoot and normally occurs in the axil of a leaf or at the tip of a stem. Once formed, a bud may remain for some time in a dormant condition, or it may form a shoot immediately. Buds may be spec ...
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Crown (botany) The crown of a plant refers to the total of an individual plant's aboveground parts, including stems, leaves, and reproductive structures. A plant community canopy consists of one or more plant crowns growing in a given area. The crown of a w ...
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Heteroblasty (botany) Heteroblasty is the significant and abrupt change in form and function, that occurs over the lifespan of certain plants. Characteristics affected include internode length and stem structure as well as leaf form, size and arrangement. It should no ...
, an abrupt change in the growth pattern of some plants as they mature *
Lateral shoot A lateral shoot, commonly known as a branch, is a part of a plant's shoot system that develops from axillary buds on the stem's surface, extending laterally from the plant's stem. Importance to photosynthesis As a plant grows it requires more ene ...
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Seedling A seedling is a young sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle (embryonic root), the hypocotyl (embry ...
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Sterigma In biology, a sterigma (pl. sterigmata) is a small supporting structure. It commonly refers to an extension of the basidium (the spore-bearing cells) consisting of a basal filamentous part and a slender projection which carries a spore at the ti ...
, the "woody peg" below the leaf of some conifers *
Thorn (botany) In plant morphology, thorns, spines, and prickles, and in general spinose structures (sometimes called ''spinose teeth'' or ''spinose apical processes''), are hard, rigid extensions or modifications of leaves, roots, stems or buds with sharp, ...
, true thorns, as distinct from spines or prickles, are short shoots


References

{{Botany Plant morphology