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The Late Cretaceous (100.5–66 Ma) is the younger of two
epochs In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided by ...
into which the Cretaceous
Period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...
is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous
Series Series may refer to: People with the name * Caroline Series (born 1951), English mathematician, daughter of George Series * George Series (1920–1995), English physicist Arts, entertainment, and media Music * Series, the ordered sets used in ...
. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as
chalk Chalk is a soft, white, porous, sedimentary carbonate rock. It is a form of limestone composed of the mineral calcite and originally formed deep under the sea by the compression of microscopic plankton that had settled to the sea floor. Chalk ...
. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period.


Climate

During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern
latitude In geography, latitude is a coordinate that specifies the north–south position of a point on the surface of the Earth or another celestial body. Latitude is given as an angle that ranges from –90° at the south pole to 90° at the north pole ...
s experienced markedly more seasonal climatic conditions.


Geography

Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves;
Appalachia Appalachia () is a cultural region in the Eastern United States that stretches from the Southern Tier of New York State to northern Alabama and Georgia. While the Appalachian Mountains stretch from Belle Isle in Newfoundland and Labrador, C ...
and
Laramidia Laramidia was an island continent that existed during the Late Cretaceous period (99.6–66 Ma), when the Western Interior Seaway split the continent of North America in two. In the Mesozoic era, Laramidia was an island land mass separated from A ...
. India maintained a northward course towards Asia. In the Southern Hemisphere, Australia and
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
seem to have remained connected and began to drift away from Africa and South America."Dinosaurs Ruled the World: Late Cretaceous Period." In: Dodson, Peter & Britt, Brooks & Carpenter, Kenneth & Forster, Catherine A. & Gillette, David D. & Norell, Mark A. & Olshevsky, George & Parrish, J. Michael & Weishampel, David B. ''The Age of Dinosaurs''. Publications International, LTD. Pp. 103-104. . Europe was an island chain. Populating some of these islands were endemic
dwarf Dwarf or dwarves may refer to: Common uses *Dwarf (folklore), a being from Germanic mythology and folklore * Dwarf, a person or animal with dwarfism Arts, entertainment, and media Fictional entities * Dwarf (''Dungeons & Dragons''), a humanoid ...
dinosaur species.


Vertebrate fauna


Non-avian dinosaurs

In the Late Cretaceous, the
hadrosaurs Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which incl ...
,
ankylosaurs Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. ...
, and
ceratopsia Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurass ...
ns experienced success in
Asiamerica Asiamerica was a supercontinent formed from the Laurasian landmass and separated by shallow continental seas from Eurasia to the West and eastern North America to the East. This region incorporated what is now politically China, Mongolia, western ...
(Western North America and eastern Asia). Tyrannosaurs dominated the large predator niche in North America. They were also present in Asia, although were usually smaller and more primitive than the North American varieties.
Pachycephalosaur Pachycephalosauria (; from Greek παχυκεφαλόσαυρος for 'thick headed lizards') is a clade of ornithischian dinosaurs. Along with Ceratopsia, it makes up the clade Marginocephalia. With the exception of two species, most pachyceph ...
s were also present in both North America and Asia.
Dromaeosaur Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning ...
s shared the same geographical distribution, and are well documented in both Mongolia and Western North America. Additionally therizinosaurs (known previously as segnosaurs) appear to have been in North America and Asia. Gondwana held a very different dinosaurian fauna, with most predators being
abelisaur Abelisauroidea is typically regarded as a Cretaceous group, though the earliest abelisauridae remains are known from the Middle Jurassic of Argentina (classified as the species Eoabelisaurus mefi) and possibly Madagascar (fragmentary remains of ...
s and carcharodontosaurs; and
titanosaur Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with taxa still thr ...
s being among the dominant herbivores.
Spinosaurids The Spinosauridae (or spinosaurids) are a clade or family of tetanuran theropod dinosaurs comprising ten to seventeen known genera. They came into prominence during the Cretaceous period. Spinosaurid fossils have been recovered worldwide, includi ...
were also present during this time.


Birds

Birds became increasingly common, diversifying in a variety of enantiornithe and
ornithurine Ornithurae (meaning "bird tails" in Greek) is a natural group which includes the common ancestor of ''Ichthyornis'', ''Hesperornis'', and all modern birds as well as all other descendants of that common ancestor. Classification Ernst Haeckel coi ...
forms. Early
Neornithes Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight ...
such as ''
Vegavis ''Vegavis'' is a genus of extinct bird that lived during the Late Cretaceous (Maastrichtian stage) of Antarctica, some 68 to 66 mya. Among modern birds, most studies show that ''Vegavis'' is most closely related to ducks and geese (Anatidae), b ...
''Supporting information
/ref> co-existed with forms as bizarre as ''
Yungavolucris ''Yungavolucris'' is a genus of enantiornithean birds. It contains the single species ''Yungavolucris brevipedalis'', which lived in the Late Cretaceous ( Maastrichtian, c.70.6 – 66 mya). The fossil bones were found in the Lecho Formation ...
'' and ''
Avisaurus ''Avisaurus'' (meaning "bird lizard") is a genus of enantiornithine bird from the Late Cretaceous of North America. Discovery ''Avisaurus archibaldi'' was discovered in the Late Cretaceous Hell Creek Formation of North America (Maastrichtian, fr ...
''. Though mostly small, marine
Hesperornithes Hesperornithes is an extinct and highly specialized group of aquatic avialans closely related to the ancestors of modern birds. They inhabited both marine and freshwater habitats in the Northern Hemisphere, and include genera such as ''Hesperorn ...
became relatively large and flightless, adapted to life in the open sea.


Pterosaurs

Though primarily represented by
azhdarchids Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cret ...
, other forms like
pteranodontids The Pteranodontidae are a family of large pterosaurs of the Cretaceous Period of North America and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a distinctive, elongated crest jutting from the rear of the head ...
, tapejarids (''
Caiuajara ''Caiuajara'' is an extinct genus of tapejarid pterosaur from the Late Cretaceous period (Santonian stage) of Brazil. It is known from a single type species, ''Caiuajara dobruskii''. Discovery and naming In 1971, the labourers Alexandre D ...
'' and ''
Bakonydraco ''Bakonydraco'' is a genus of pterodactyloid pterosaur from the Late Cretaceous period ( Santonian stage) of what is now the Csehbánya Formation of the Bakony Mountains, Iharkút, Veszprém, western Hungary. Etymology ''Bakonydraco'' was n ...
''),
nyctosaurids Nyctosauridae (meaning "night lizards" or "bat lizards") is a family of specialized soaring pterosaurs of the late Cretaceous Period of North America, Africa, and possibly Europe. It was named in 1889 by Henry Alleyne Nicholson and Richard Lyde ...
and uncertain forms (''
Piksi ''Piksi'' is a potential pterosaurLongrich, N.R., Martill, D.M., and Andres, B. (2018). Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary. PLoS Biology, 16(3): e2001663. genu ...
'', ''
Navajodactylus ''Navajodactylus'' (meaning "Navajo finger") is an extinct genus of pterodactyloid pterosaur from Late Cretaceous (late Campanian stage) deposits of the San Juan Basin in New Mexico, United States. Discovery The holotype specimen of ''Navajodac ...
'') are also present. Historically, it has been assumed that pterosaurs were in decline due to competition with birds, but it appears that neither group overlapped significantly ecologically, nor is it particularly evident that a true systematic decline was ever in place, especially with the discovery of smaller pterosaur species.


Mammals

Several old mammal groups began to disappear, with the last eutriconodonts occurring in the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian ...
of North America. In the northern hemisphere,
cimolodont Cimolodonta is a taxon of extinct mammals that lived from the Cretaceous to the Eocene. They were some of the more derived members of the extinct order Multituberculata. They probably lived something of a rodent-like existence until their ecolo ...
,
multituberculates Multituberculata (commonly known as multituberculates, named for the multiple tubercles of their teeth) is an extinct order of rodent-like mammals with a fossil record spanning over 130 million years. They first appeared in the Middle Jurassic, ...
, metatherians and
eutherian Eutheria (; from Greek , 'good, right' and , 'beast'; ) is the clade consisting of all therian mammals that are more closely related to placentals than to marsupials. Eutherians are distinguished from noneutherians by various phenotypic tra ...
s were the dominant mammals, with the former two groups being the most common mammals in North America. In the southern hemisphere there was instead a more complex fauna of
dryolestoid Dryolestida is an extinct order of mammals, primarily and possibly exclusively known from the Jurassic and Cretaceous. They are considered members of the clade Cladotheria, close to the ancestry of therian mammals. It is also believed that they d ...
s,
gondwanathere Gondwanatheria is an extinct group of mammaliaforms that lived in parts of Gondwana, including Madagascar, India, South America, Africa and Antarctica during the Upper Cretaceous through the Paleogene (and possibly much earlier, if '' Allostaffia ...
s and other multituberculates and basal
eutherian Eutheria (; from Greek , 'good, right' and , 'beast'; ) is the clade consisting of all therian mammals that are more closely related to placentals than to marsupials. Eutherians are distinguished from noneutherians by various phenotypic tra ...
s; monotremes were presumably present, as was the last of the
haramiyida Haramiyida ("thief" from Arabic الحرامية (al ḥarāmiyah), "thief, bandit") is a possibly polyphyletic order of mammaliaform cynodonts or mammals of controversial taxonomic affinites. Their teeth, which are by far the most common remains ...
ns, ''Avashishta''. Mammals, though generally small, ranged into a variety of ecological niches, from carnivores (
Deltatheroida Deltatheroida is an extinct group of basal metatherians that were distantly related to modern marsupials. The majority of known members of the group lived in the Cretaceous; one species, '' Gurbanodelta kara'', is known from the late Paleocene ( ...
), to mollusc-eater (
Stagodontidae Stagodontidae is an extinct family of carnivorous metatherian mammals that inhabited North America and Europe during the late Cretaceous, and possibly to the Eocene in South America. Description Currently, the family includes four genera, ' ...
), to herbivores (multituberculates, ''
Schowalteria ''Schowalteria'' is a genus of extinct mammal from the Cretaceous of Canada. It is the earliest known representative of Taeniodonta, a specialised lineage of non-placental eutherian mammals otherwise found in Paleocene and Eocene deposits. It is ...
'',
Zhelestidae Zhelestidae is a lineage of extinct eutherian mammals. Occurring in the Late Cretaceous from the Turonian to the Maastrichtian, they were an extremely successful group, with representatives present in Europe, Asia, India (and subsequently in Ma ...
and Mesungulatidae) to highly atypical cursorial forms (
Zalambdalestidae Zalambdalestidae is a clade of Asian eutherians occurring during the Late Cretaceous. Once classified as Glires, features like epipubic bones and various cranial elements have identified these animals as outside of Placentalia, representing thus ...
,
Brandoniidae Dryolestidae is an extinct family of Mesozoic mammals, known from the Middle Jurassic to the Early Cretaceous of the North Hemisphere. The oldest known member, '' Anthracolestes'', is known from the Middle Jurassic Itat Formation of Western Sib ...
). True placentals evolved only at the very end of the epoch; the same can be said for true marsupials. Instead, nearly all known eutherian and metatherian fossils belong to other groups.


Marine life

In the seas, mosasaurs suddenly appeared and underwent a spectacular evolutionary radiation. Modern sharks also appeared and giant-penguin-like polycotylid plesiosaurs (3 meters long) and huge long-necked elasmosaurs (13 meters long) also diversified. These predators fed on the numerous teleost fishes, which in turn evolved into new advanced and modern forms ( Neoteleostei). Ichthyosaurs and
pliosaur Pliosauroidea is an extinct clade of plesiosaurs, known from the earliest Jurassic to early Late Cretaceous. They are best known for the subclade Thalassophonea, which contained crocodile-like short-necked forms with large heads and massive toot ...
s, on the other hand, became extinct during the
Cenomanian-Turonian anoxic event The Cenomanian-Turonian boundary event, also known as the Cenomanian-Turonian extinction, Cenomanian-Turonian oceanic anoxic event ( OAE 2), and referred to also as the Bonarelli event, was one of two anoxic extinction events in the Cretaceous p ...
.


Flora

Near the end of the Cretaceous Period, flowering plants diversified. In temperate regions, familiar plants like
magnolia ''Magnolia'' is a large genus of about 210 to 340The number of species in the genus ''Magnolia'' depends on the taxonomic view that one takes up. Recent molecular and morphological research shows that former genera ''Talauma'', ''Dugandiodendr ...
s, sassafras, roses,
redwood Sequoioideae, popularly known as redwoods, is a subfamily of coniferous trees within the family Cupressaceae. It includes the largest and tallest trees in the world. Description The three redwood subfamily genera are '' Sequoia'' from coasta ...
s, and willows could be found in abundance.


Cretaceous–Paleogene mass extinction discovery

The Cretaceous–Paleogene extinction event was a large-scale
mass extinction An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp change in the diversity and abundance of multicellular organisms. It ...
of animal and plant species in a geologically short period of time, approximately (Ma). It is widely known as the ''K–T extinction event'' and is associated with a geological signature, usually a thin band dated to that time and found in various parts of the world, known as the
Cretaceous–Paleogene boundary The Cretaceous–Paleogene (K–Pg) boundary, formerly known as the Cretaceous–Tertiary (K–T) boundary, is a geological signature, usually a thin band of rock containing much more iridium than other bands. The K–Pg boundary marks the end of ...
(K–T boundary). ''K'' is the traditional abbreviation for the Cretaceous Period derived from the German name ''Kreidezeit'', and ''T'' is the abbreviation for the
Tertiary Tertiary ( ) is a widely used but obsolete term for the geologic period from 66 million to 2.6 million years ago. The period began with the demise of the non-avian dinosaurs in the Cretaceous–Paleogene extinction event, at the start ...
Period (a historical term for the period of time now covered by the Paleogene and Neogene periods). The event marks the end of the Mesozoic Era and the beginning of the
Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configurat ...
Era. "Tertiary" being no longer recognized as a formal time or rock unit by the International Commission on Stratigraphy, the K-T event is now called the Cretaceous—Paleogene (or K-Pg) extinction event by many researchers. Non-
avian Avian may refer to: * Birds or Aves, winged animals *Avian (given name) (russian: Авиа́н, link=no), a male forename Aviation *Avro Avian, a series of light aircraft made by Avro in the 1920s and 1930s *Avian Limited, a hang glider manufactur ...
dinosaur
fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...
are found only below the Cretaceous–Paleogene boundary and became extinct immediately before or during the event. A very small number of dinosaur fossils have been found above the Cretaceous–Paleogene boundary, but they have been explained as reworked fossils, that is, fossils that have been eroded from their original locations then preserved in later sedimentary layers. Mosasaurs, plesiosaurs,
pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...
s and many
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
of plants and invertebrates also became extinct.
Mammalian Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur o ...
and bird clades passed through the boundary with few extinctions, and
evolutionary radiation An evolutionary radiation is an increase in taxonomic diversity that is caused by elevated rates of speciation, that may or may not be associated with an increase in morphological disparity. Radiations may affect one clade or many, and be rapid ...
from those Maastrichtian clades occurred well past the boundary. Rates of extinction and radiation varied across different clades of organisms. Many scientists hypothesize that the Cretaceous–Paleogene extinctions were caused by catastrophic events such as the massive
asteroid impact An impact event is a collision between astronomical objects causing measurable effects. Impact events have physical consequences and have been found to regularly occur in planetary systems, though the most frequent involve asteroids, comets or ...
that caused the
Chicxulub crater The Chicxulub crater () is an impact crater buried underneath the Yucatán Peninsula in Mexico. Its center is offshore near the community of Chicxulub, after which it is named. It was formed slightly over 66 million years ago when a large as ...
, in combination with increased
volcanic activity Volcanism, vulcanism or volcanicity is the phenomenon of eruption of molten rock (magma) onto the surface of the Earth or a solid-surface planet or moon, where lava, pyroclastics, and volcanic gases erupt through a break in the surface called a ...
, such as that recorded in the Deccan Traps, both of which have been firmly dated to the time of the extinction event. In theory, these events reduced sunlight and hindered photosynthesis, leading to a massive disruption in Earth's ecology. A much smaller number of researchers believe the extinction was more gradual, resulting from slower changes in sea level or
climate Climate is the long-term weather pattern in an area, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorologi ...
.


See also

*
Flora and fauna of the Maastrichtian stage This is an incomplete list that briefly describes vertebrates that were extant during the Maastrichtian, a stage of the Late Cretaceous Period which extended from 72.1 to 66 million years before present. This was the last time period in which ...
*
Hațeg Island Hațeg Island was a large offshore island in the Tethys Sea which existed during the Late Cretaceous period, probably from the Cenomanian to the Maastrichtian ages. It was situated in an area corresponding to the region around modern-day Hațeg, ...


References

{{Geological history, p, m Late Geological epochs