Sematuridae is a

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...

s in the lepidopteran order that contains two subfamilies ( Minet and Scoble, 1999).

Taxonomy, systematics, and identification

These are large day- or night-flying moths, usually tailed and similar to

Uraniidae The Uraniidae are a family of moths containing four subfamilies, 90 genera, and roughly 700 species. The family is distributed throughout the tropics of the Americas, Africa and Indo-Australia.Carter, David, ''Eyewitness Handbook to Butterflies ...

in general appearanc

(except for the genera ''Apoprogones'', ''Anuropteryx'' and ''Lonchotura''). The position of this family is not certain amongst the

Macrolepidoptera Macrolepidoptera is a group within the insect order Lepidoptera. Traditionally used for the larger butterflies and moths as opposed to the "microlepidoptera", this group is artificial. However, it seems that by moving some taxa about, a monophy ...

but it is usually considered to belong to the superfamily

Geometroidea The Geometroidea are the superfamily of geometrid moths in the order Lepidoptera. It includes the families Geometridae, Uraniidae, Epicopeiidae, Sematuridae, and the recently established family Pseudobistonidae. The monotypic genus ''Apoprogones ...

, whilst the identity of its closest extant relative is under investigation using DNA sequencing

Until recently very little has been known of sematurid biology (but see

Sematurinae Sematurinae is a subfamily of moths in the family Sematuridae represented by at least 29 species in the Neotropics. Taxonomy, classification and identification Sematurinae has three principal genera, and possibly another two. The hindwing of ''C ...

). The "chaetosemata" of Sematuridae have long

setae In biology, setae (singular seta ; from the Latin word for " bristle") are any of a number of different bristle- or hair-like structures on living organisms. Animal setae Protostomes Annelid setae are stiff bristles present on the body. ...

overhanging the

compound eyes A compound eye is a visual organ found in arthropods such as insects and crustaceans. It may consist of thousands of ommatidia, which are tiny independent photoreception units that consist of a cornea, lens, and photoreceptor cells which distin ...

, the antennae are thickened towards the often rather

clubbed ''Clubbed'' is a 2008 British drama film about a 1980s factory worker who takes up a job as a club doorman, written by Geoff Thompson and directed by Neil Thompson. Plot In 1984, Danny - a lonely factory worker intimidated by life - is batter ...

or hooked tip whilst a

tympanal organ A tympanal organ (or tympanic organ) is a hearing organ in insects, consisting of a membrane ( tympanum) stretched across a frame backed by an air sac and associated sensory neurons. Sounds vibrate the membrane, and the vibrations are sensed by ...

for hearing is always absent and there are other diagnostic characters in the

abdomen The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the to ...

(Minet and Scoble, 1999: 305-307; Holloway et al., 2001).

Distribution

Apoprogoninae is represented by a single species in

South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...

whilst

is represented by about 29 (41 including the genera ''Anurapteryx'' and ''Lonchotura'') species in the

Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...

. Such an apparently relictual distribution might relate to the geological split of

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...

and

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

, but there is as yet no evidence for the age of

evolutionary Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...

divergence of

Neotropical The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bioge ...

and Afrotropical representatives; moreover, new genetic material would be needed for the

taxon In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...

as well as the

American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...

genera ''Anurapteryx'' and ''Lonchotura'' for a modern molecular approach to this problem to succee

References

*Holloway, J.D., Kibby, G and Peggie, D. (1997). ''The families of Malesian moths and butterflies''. Fauna Malesia Handbooks. 455 pp. Brill Academic Publishers, Leiden. *Minet, J. and Scoble, M. J. (1999)

998 Year 998 ( CMXCVIII) was a common year starting on Saturday (link will display the full calendar) of the Julian calendar. Events By place Europe * Spring – Otto III retakes Rome and restores power in the papal city. Crescenti ...

The Drepanoid/Geometroid Assemblage. Pp. 301–320 in Kristensen, N.P. (Ed.), ''Lepidoptera, Moths and Butterflies. Volume 1, Evolution, Systematics, and Biogeography''. Handbook of Zoology, vol. IV, Arthropoda: Insecta, Part 35: 491 pp. Walter de Gruyter, Berlin and New York. {{Taxonbar, from=Q1935965 Moth families