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Seaweed, or macroalgae, refers to thousands of species of
macroscopic The macroscopic scale is the length scale on which objects or phenomena are large enough to be visible with the naked eye, without magnifying optical instruments. It is the opposite of microscopic. Overview When applied to physical phenomena a ...
,
multicellular A multicellular organism is an organism that consists of more than one cell, in contrast to unicellular organism. All species of animals, land plants and most fungi are multicellular, as are many algae, whereas a few organisms are partially uni ...
, marine
algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthesis, photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from u ...
. The term includes some types of '' Rhodophyta'' (red), ''
Phaeophyta Brown algae (singular: alga), comprising the class Phaeophyceae, are a large group of multicellular algae, including many seaweeds located in colder waters within the Northern Hemisphere. Brown algae are the major seaweeds of the temperate and ...
'' (brown) and ''
Chlorophyta Chlorophyta or Prasinophyta is a taxon of green algae informally called chlorophytes. The name is used in two very different senses, so care is needed to determine the use by a particular author. In older classification systems, it refers to a ...
'' (green) macroalgae. Seaweed species such as
kelp Kelps are large brown algae seaweeds that make up the order Laminariales. There are about 30 different genera. Despite its appearance, kelp is not a plant - it is a heterokont, a completely unrelated group of organisms. Kelp grows in "underwat ...
s provide essential nursery habitat for fisheries and other marine species and thus protect food sources; other species, such as planktonic algae, play a vital role in capturing carbon, producing at least 50% of Earth's oxygen. Natural seaweed ecosystems are sometimes under threat from human activity. For example, mechanical
dredging Dredging is the excavation of material from a water environment. Possible reasons for dredging include improving existing water features; reshaping land and water features to alter drainage, navigability, and commercial use; constructing da ...
of kelp destroys the resource and dependent fisheries. Other forces also threaten some seaweed ecosystems; a wasting disease in predators of purple urchins has led to a urchin population surge which destroyed large kelp forest regions off the coast of California. Humans have a long history of cultivating seaweeds for their uses. In recent years,
seaweed farming Seaweed farming or kelp farming is the practice of cultivating and harvesting seaweed. In its simplest form, it consists of the management of naturally found batches. In its most advanced form, it consists of fully controlling the life cycle ...
has become a global agricultural practice, providing food, source material for various chemical uses (such as
carrageenan Carrageenans or carrageenins ( ; ) are a family of natural linear sulfated polysaccharides that are extracted from red edible seaweeds. Carrageenans are widely used in the food industry, for their gelling, thickening, and stabilizing properties. ...
), cattle feeds and fertilizers. Because of their importance in marine ecologies and for absorbing carbon dioxide, recent attention has been on cultivating seaweeds as a potential
climate change mitigation Climate change mitigation is action to limit climate change by reducing emissions of greenhouse gases or removing those gases from the atmosphere. The recent rise in global average temperature is mostly caused by emissions from fossil fuels bu ...
strategy for biosequestration of carbon dioxide, alongside other benefits like nutrient pollution reduction, increased habitat for coastal aquatic species, and reducing local
ocean acidification Ocean acidification is the reduction in the pH value of the Earth’s ocean. Between 1751 and 2021, the average pH value of the ocean surface has decreased from approximately 8.25 to 8.14. The root cause of ocean acidification is carbon dioxide ...
. The
IPCC The Intergovernmental Panel on Climate Change (IPCC) is an intergovernmental body of the United Nations. Its job is to advance scientific knowledge about climate change caused by human activities. The World Meteorological Organization (WMO) ...
Special Report on the Ocean and Cryosphere in a Changing Climate recommends "further research attention" as a mitigation tactic.


Taxonomy

"Seaweed" lacks a formal definition, but seaweed generally lives in the ocean and is visible to the naked eye. The term refers to both flowering plants submerged in the ocean, like eelgrass, as well as larger marine algae. Generally it is one of several groups of multicellular
algae Algae (; singular alga ) is an informal term for a large and diverse group of photosynthesis, photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from u ...
:
red Red is the color at the long wavelength end of the visible spectrum of light, next to orange and opposite violet. It has a dominant wavelength of approximately 625–740 nanometres. It is a primary color in the RGB color model and a secondary ...
, green and
brown Brown is a color. It can be considered a composite color, but it is mainly a darker shade of orange. In the CMYK color model used in printing or painting, brown is usually made by combining the colors orange and black. In the RGB color model used ...
. They lack one common multicellular ancestor, forming a polyphyletic group. In addition, bluegreen algae ( Cyanobacteria) are occasionally considered in seaweed literature. The number of seaweed species is still discussed among scientists, but most likely there are several thousand species of seaweed.


Genera

The following table lists a very few example genera of seaweed.


Anatomy

Seaweed's appearance resembles non- woody terrestrial plants. Its anatomy includes: * Thallus: algal body ** Lamina or blade: flattened structure that is somewhat leaf-like ***
Sorus A sorus (pl. sori) is a cluster of sporangia (structures producing and containing spores) in ferns and fungi. A coenosorus (plural coenosori) is a compound sorus composed of multiple, fused sori. Etymology This New Latin word is from Ancient G ...
:
spore In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, ...
cluster *** ''
pneumatocyst In phycology, a pneumatocyst is a floating structure that contains gas found on brown seaweed. A seaweed's thallus may have more than one. They provide buoyancy Buoyancy (), or upthrust, is an upward force exerted by a fluid that oppose ...
'', air bladder: a flotation-assisting organ on the blade ***
Kelp Kelps are large brown algae seaweeds that make up the order Laminariales. There are about 30 different genera. Despite its appearance, kelp is not a plant - it is a heterokont, a completely unrelated group of organisms. Kelp grows in "underwat ...
, float: a flotation-assisting organ between the lamina and stipe ** Stipe: stem-like structure, may be absent ** Holdfast: basal structure providing attachment to a substrate ***
Haptera A holdfast is a root-like structure that anchors aquatic sessile organisms, such as seaweed, other sessile algae, stalked crinoids, benthic cnidarians, and sponges, to the substrate. Holdfasts vary in shape and form depending on both the spec ...
: finger-like extension of the holdfast that anchors to a benthic substrate The stipe and blade are collectively known as the
frond A frond is a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group. Other botanists allow the term frond to also apply to the lar ...
.


Ecology

Two environmental requirements dominate seaweed ecology. These are seawater (or at least
brackish water Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
) and light sufficient to support photosynthesis. Another common requirement is an attachment point, and therefore seaweed most commonly inhabits the littoral zone (nearshore waters) and within that zone, on rocky shores more than on sand or shingle. In addition, there are few genera (e.g., '' Sargassum'' and ''
Gracilaria ''Gracilaria'' is a genus of red algae (Rhodophyta) notable for its economic importance as an agarophyte, as well as its use as a food for humans and various species of shellfish. Various species within the genus are cultivated among Asia, Sou ...
'') which do not live attached to the sea floor, but float freely. Seaweed occupies various ecological niches. At the surface, they are only wetted by the tops of sea spray, while some species may attach to a substrate several meters deep. In some areas, littoral seaweed colonies can extend miles out to sea. The deepest living seaweed are some species of red algae. Others have adapted to live in tidal rock pools. In this habitat, seaweed must withstand rapidly changing temperature and salinity and occasional drying. Macroalgae and macroalgal detritus have also been shown to be an important food source for benthic organisms, because macroalgae shed old fronds.. These macroalgal fronds tend to be utilized by
benthos Benthos (), also known as benthon, is the community of organisms that live on, in, or near the bottom of a sea, river, lake, or stream, also known as the benthic zone.intertidal zone The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species of ...
close to the shore. Alternatively, pneumatocysts (gas filled “bubbles”) can keep the macroalgae thallus afloat fronds are transported by wind and currents from the coast into the deep ocean. It has been shown that
benthic organisms Benthos (), also known as benthon, is the community of organisms that live on, in, or near the bottom of a sea, river, lake, or stream, also known as the benthic zone. As macroalgae takes up carbon dioxide and release oxygen in the photosynthesis, macroalgae fronds can also contribute to
carbon sequestration Carbon sequestration is the process of storing carbon in a carbon pool. Carbon dioxide () is naturally captured from the atmosphere through biological, chemical, and physical processes. These changes can be accelerated through changes in lan ...
in the ocean, when the macroalgal fronds drift offshore into the deep
ocean basins In hydrology, an oceanic basin (or ocean basin) is anywhere on Earth that is covered by seawater. Geologically, ocean basins are large  geologic basins that are below sea level. Most commonly the ocean is divided into basins foll ...
and sink to the sea floor without being remineralized by organisms. The importance of this process for the
Blue Carbon Blue Carbon refers to organic carbon that is captured and stored by the world's oceanic and coastal ecosystems, mostly by algae, seagrasses, macroalgae, mangroves, salt marshes and other plants in coastal wetlands. The term Blue Carbon was coine ...
storage is currently discussed among scientists.


Biogeographic Expansion

Nowadays a number of vectors - e.g., transport on ship hulls, exchanges among shellfish farmers, global warming, opening of trans-oceanic canals - all combine to enhance the transfer of exotic seaweeds to new environments. Since the piercing of the Suez Canal,the situation is particularly acute in the Mediterranean Sea, a 'marine biodiversity hotspot' that now registers over 120 newly introduced seaweed species -the largest number in the world.


Production

As of 2018, the top 10 countries produced 96% of the global total of 2,165,675 metric tons.


Farming


Uses

Seaweed has a variety of uses, for which it is
farmed Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people to ...
or foraged.


Food

Seaweed is consumed across the world, particularly in East Asia, e.g. Japan, China, Korea, Taiwan and Southeast Asia, e.g. Brunei, Singapore, Thailand,
Burma Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
,
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand ...
, Vietnam, Indonesia, the Philippines, and Malaysia, as well as in South Africa,
Belize Belize (; bzj, Bileez) is a Caribbean and Central American country on the northeastern coast of Central America. It is bordered by Mexico to the north, the Caribbean Sea to the east, and Guatemala to the west and south. It also shares a wate ...
, Peru,
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
, the
Canadian Maritimes The Maritimes, also called the Maritime provinces, is a region of Eastern Canada consisting of three provinces: New Brunswick, Nova Scotia, and Prince Edward Island. The Maritimes had a population of 1,899,324 in 2021, which makes up 5.1% of Ca ...
, Scandinavia,
South West England South West England, or the South West of England, is one of nine official regions of England. It consists of the counties of Bristol, Cornwall (including the Isles of Scilly), Dorset, Devon, Gloucestershire, Somerset and Wiltshire. Cities and ...
, Ireland, Wales, Hawaii and California, and Scotland. Gim (김, Korea),
nori Nori is a dried edible seaweed used in Japanese cuisine, made from species of the red algae genus '' Pyropia'', including ''P. yezonesis'' and '' P. tenera''. It has a strong and distinctive flavor, and is often used to wrap rolls of sushi or ...
(海苔, Japan) and zicai (紫菜, China) are sheets of dried ''
Porphyra ''Porphyra'' is a genus of coldwater seaweeds that grow in cold, shallow seawater. More specifically, it belongs to red algae phylum of laver species (from which comes laverbread), comprising approximately 70 species.Brodie, J.A. and Irvine, ...
'' used in soups,
sushi is a Japanese dish of prepared , usually with some sugar and salt, accompanied by a variety of , such as seafood, often raw, and vegetables. Styles of sushi and its presentation vary widely, but the one key ingredient is "sushi rice," also ...
or ''
onigiri , also known as , , or rice ball, is a Japanese food made from white rice formed into triangular or cylindrical shapes and often wrapped in ''nori''. Traditionally, an onigiri is filled with pickled ume ('' umeboshi''), salted salmon, katsuob ...
'' (rice balls). ''
Chondrus crispus ''Chondrus crispus''—commonly called Irish moss or carrageen moss (Irish ''carraigín'', "little rock")—is a species of red algae which grows abundantly along the rocky parts of the Atlantic coast of Europe and North America. In its fres ...
'' ('Irish moss' or
carrageenan Carrageenans or carrageenins ( ; ) are a family of natural linear sulfated polysaccharides that are extracted from red edible seaweeds. Carrageenans are widely used in the food industry, for their gelling, thickening, and stabilizing properties. ...
moss) is used in food additives, along with '' Kappaphycus'' and G''igartinoid'' seaweed. ''Porphyra'' is used in Wales to make
laverbread Laverbread (; cy, bara lafwr or '; ga, sleabhac) is a food product made from laver, an edible seaweed (littoral alga) consumed mainly in Wales as part of local traditional cuisine. The seaweed is commonly found around the west coast of Great ...
(sometimes with
oat The oat (''Avena sativa''), sometimes called the common oat, is a species of cereal grain grown for its seed, which is known by the same name (usually in the plural, unlike other cereals and pseudocereals). While oats are suitable for human con ...
flour). In northern
Belize Belize (; bzj, Bileez) is a Caribbean and Central American country on the northeastern coast of Central America. It is bordered by Mexico to the north, the Caribbean Sea to the east, and Guatemala to the west and south. It also shares a wate ...
, seaweed is mixed with milk, nutmeg,
cinnamon Cinnamon is a spice obtained from the inner bark of several tree species from the genus '' Cinnamomum''. Cinnamon is used mainly as an aromatic condiment and flavouring additive in a wide variety of cuisines, sweet and savoury dishes, break ...
and vanilla to make "" ("sweet").
Alginate Alginic acid, also called algin, is a naturally occurring, edible polysaccharide found in brown algae. It is hydrophilic and forms a viscous gum when hydrated. With metals such as sodium and calcium, its salts are known as alginates. Its colour ...
,
agar Agar ( or ), or agar-agar, is a jelly-like substance consisting of polysaccharides obtained from the cell walls of some species of red algae, primarily from ogonori ('' Gracilaria'') and "tengusa" ('' Gelidiaceae''). As found in nature, agar ...
and
carrageenan Carrageenans or carrageenins ( ; ) are a family of natural linear sulfated polysaccharides that are extracted from red edible seaweeds. Carrageenans are widely used in the food industry, for their gelling, thickening, and stabilizing properties. ...
are gelatinous seaweed products collectively known as
hydrocolloid A colloid is a mixture in which one substance consisting of microscopically dispersed insoluble particles is suspended throughout another substance. Some definitions specify that the particles must be dispersed in a liquid, while others extend ...
s or
phycocolloid {{Short pages monitor