Sciurumimus

''Sciurumimus'' ("Squirrel-mimic," named for its tail's resemblance to that of the tree squirrel, ''Sciurus'') is an extinct genus of tetanuran theropod from the Late Jurassic of Germany. It is known from a single juvenile specimen representing the type species, ''Sciurumimus albersdoerferi'', which was found in a limestone quarry close to Painten in Lower Bavaria. The specimen was preserved with traces of feather-like filaments.

The ''Sciurumimus'' specimen was first announced in 2011 in an informal presentation, but not formally described and named until the following year. Although originally classified as a basal megalosauroid, later phylogenetic analyses dispute this placement, suggesting it was a basal member of the Orionides.

Description

''Sciurumimus'' is known from a single holotype fossil that is exceptionally well-preserved, with full skeleton in complete articulation along with fine details of soft tissue. It is comparable in size and proportions to the juvenile coelurosaur ''Juravenator'', although differs significantly in several anatomical details. The skull of ''Sciurumimus'' is proportionally large, at 156% of the length of the femur and longer than the cervical vertebrae series. These body proportions, along with short forelimbs, lack of fusion in the skeleton, and regular tooth morphology indicate the specimen represents a very young, probably early-posthatchling individual.

The fossil preserves filamentous plumage at the tail base and on other parts of the body. These structures are described as being identical to the stage 1 feathers preserved in some ornithischians, the basal tyrannosaur ''Dilong'', and the basal therizinosauroid '' Beipiaosaurus''. Skin is also preserved. Although most of the preserved soft tissue on the ''Sciurumimus'' holotype likely represent integumentary structures, a small patch of what may be muscle tissue is observed along the rear edge of the tibia.

Classification

When first discovered, a phylogenetic analysis suggested that ''Sciurumimus'' may have been a primitive member of the Megalosauroidea, a clade of large carnivorous dinosaurs more primitive than many other well-known theropods like the tyrannosauroids and carnosaurs, making it the most basal known feathered theropod. This classification was supported by one of three initial analyses conducted by the scientists who described it. The other two analyses, which the authors regarded as less well supported, found it as more closely related to ''Monolophosaurus'' and Avetheropoda, and in an unresolved position among avetheropods and megalosauroids, respectively. The exact position in the various analyses was difficult to determine due to the fact that the only known specimen is a very young juvenile.

This initial study was criticized by several researchers, who noted that some of the old analysis the scientists used to plug in data from the new fossil were incomplete and missing relevant data on various species.Mortimer, M. 2013.
Coelurosauria
", The Theropod Database. Accessed online 09-AUG-2013.http://archosaur.us/theropoddatabase/Coelurosauria.htm#SciurumimusalbersdoerferiMortimer, M. and Marjanovic, D. (in prep.) http://archosaur.us/theropoddatabase/Coelurosauria.htm#Sciurumimusalbersdoerferi In an analysis published in 2013 ''Sciurumimus'' was obtained as one of the most primitive members of the Coelurosauria, more derived than the megalosauroids. However, this analysis focused on the Coelurosauria and the dataset on which it was based contained only one megalosaurid and only four terminal taxa outside the Coelurosauria.See pages 3 and 11 in th
supplementary material
of Other analyses found it to be an allosauroid or a tetanuran outside Avetheropoda. Paul (2016) placed it among basal neocoelurosaurs. A detailed reanalysis of the taxon was done in the supplementary material of Hartman ''et al''. (2019) which found several characters to be misscored, and when the uncorrected ''Sciurumimus'' was placed in the matrix ''Juravenator'' was also found to be a basal tetanuran, while if ''Sciurumimus'' was absent ''Juravenator'' would be found to be a basal coelurosaur. The analysis found ''Sciurumimus'' to be a compsognathid, though they noted that this position was tentative. Foth ''et al''. (2020) again placed, albeit tentatively, ''Sciurumimus'' in Megalosauroidea, this time accompanied by ''Juravenator''.

References

{{Taxonbar, from=Q133894

Prehistoric coelurosaurs
Late Jurassic dinosaurs of Europe
Fossils of Germany
Feathered dinosaurs
Fossil taxa described in 2012
Taxa named by Mark Norell